List of Gene Names and Symbols
GeneAlleleLocus NameAllele NameSynonyms of AllelePhenotype
(K)--   Inhibits modifiers; permitting expression of tangerine gene; interacts with yellow fruit flesh gene.
a--anthocyaninless a1Stems and leaves always lack anthocyanin.
aprov2anthocyaninlessprovisional 2aStems and leaves always lack anthocyanin.
aprov3anthocyaninlessprovisional 3aStems and leaves always lack anthocyanin.
aa--anthocyanin absent  Complete absence of anthocyanin in all parts
Abg--Aubergine  Fruit epidermis purple, particularly on shoulder and where exposed to direct light; also enhanced by wounding.
abi--aborted inflorescence  Inflorescences reduced to an abortive 2-3 mm structure.
ac--apocarpous  Highly deformed multicarpellate and apocarpous fruits
Aco-11Aconitase-1   
Aco-12Aconitase-1   
Aco-13Aconitase-1   
Aco-22Aconitase-2   
acr--acroxantha acr1Smaller plant; short internodes; rugose, dull, light green leaves, turn yellow starting at tip.
acu-2--acumbens-2 acu2Short, rounded cotyledons; short, compacted leaves.
ad--Alternaria alternata resistance  Resistance to Alternaria collar rot.
Adh-11Alcohol dehydrogenase-1   
Adh-12Alcohol dehydrogenase-1   
Adh-1nAlcohol dehydrogenase-1   
Adh-21Alcohol dehydrogenase-2   
adp--adpressa  Plant small and decumbent with short internodes and leaves.
adp-2--adpressa-2 adp2Prostrate plant habit; short internodes and leaves; rugose, bright green pinnae.
adu--adusta adu1Small plants, proportionately reduced; progressive necrosis from older to younger leaves.
adu-2--adusta-2 adu2Stems and leaves shortened, develop progressive necrosis; small, rounded, rugose pinnae
adu-3--adusta-3 adu3Slightly smaller leaves, shiny above, develop progressive necrosis.
ae--entirely anthocyaninless a332Completely free of anthocyanin.
ae2entirely anthocyaninless  Completely free of anthocyanin.
aeafrentirely anthocyaninlessfragilisafr, apAnthocyanin absent; small, brittle, fragile plants; leaves wilt in daytime.
aeprov3entirely anthocyaninlessprovisional 3aeLacks anthocyanin in the seedling stem.
aeg--aegrota  Brown necrotic layer spreads over leaves, which die prematurely.
aeg-2--aegrota-2 aeg2Dainty leaves turn yellow and die early.
aer--aerial roots  Adventitious roots on stem from soil level to considerable height above.
aer-2--aerial roots-2  Abundance of root initials along the stems.
af--anthocyanin free a325Completely free of anthocyanin; lacks glandular hairs.
afe--afertilis afe1Small plant; short internodes, leaves; dainty, yellowish, mostly keeled pinnae; poor fruit set; heterozygote intermediate.
afl--albifolium afSeedling very slow; cotyledons white; leaves becoming normal except for gray flecks; later growth nearly normal.
afl-2--albifolium-2  Cotyledons whitish light green; pinnae with irregular yellow-white areas; green-white stripes on the stems and petioles; foliage of mature plant grey-green with whitish blotches; plants erect, 1/3 - 2/5 normal size. Slightly dominant, the heterozygote attaining 2/3 -3/4 normal size.
Aft--Anthocyanin fruit AfAnthocyanin in green and ripe fruit; environmentally sensitive, absent when shaded
ag--anthocyanin gainer  Anthocyanin absent except on cotyledons and lower sides of leaves; anthocyanin accumulation more intense under growth retarding conditions; under low temperature and low light intensity, foliage becomes yellowish, webs of corolla-like tissue appear between segments of the calyx, and fertility may be reduced.
ag(1k)anthocyanin gainerKnowlesag^3Anthocyanin absent except on cotyledons and lower sides of leaves. (Indistinguishable from ag).
ag(1s)anthocyanin gainerStubbe'sag^2Anthocyanin absent except on cotyledons and lower sides of leaves.
ag(2k)anthocyanin gainerKnowles 2ndag^4Anthocyanin absent except on cotyledons and lower sides of leaves.
ag2anthocyanin gainer  Completely free of anthocyanin.
agkanthocyanin gainerKosala's As for ag, purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
agsanthocyanin gainerSolanum Typical ag phenotype.
ag-2--anthocyanin gainer-2  As for ag purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
ags-2--aegrescens-2  Plant 1/3 normal size; reduced branching; leaves shortened; pinnae small, olive green, becoming necrotic at tips.
ah--Hoffman's anthocyaninless ao, a337Completely free of anthocyanin
ahprov2Hoffman's anthocyaninlessprovisional 2ahCompletely free of anthocyanin
ahprov3Hoffman's anthocyaninlessprovisional 3ahCompletely free of anthocyanin
ahprov4Hoffman's anthocyaninlessprovisional 4ahCompletely free of anthocyanin
ahprov5Hoffman's anthocyaninlessprovisional 5ahCompletely free of anthocyanin
ahprov6Hoffman's anthocyaninlessprovisional 6ahCompletely free of anthocyanin
ahprov7Hoffman's anthocyaninlessprovisional 7ahCompletely free of anthocyanin
ai--incomplete anthocyanin a342Early seedling stem has trace of anthocyanin, soon lost.
ai2incomplete anthocyanin am, a340Early seedling stem has trace of anthocyanin, soon lost. (Typical green stem mutant.)
al--anthocyanin loser a2Anthocyanin pigmentation of medium intensity appears for ten to twenty days after seedling emergence, thereafter disappearing except for islands near the leaf nodes; within three days after this process begins, the stem becomes entirely green and remains so for the rest of the life of the plant..
alpu-2anthocyanin loserpulvinata-2pu-2Semispherical bush, anthocyaninless; short internodes.
al-2--anthocyanin loser-2  No anthocyanin in cotyledons and primary leaves; little anthocyanin in hypocotyl; internodes and leaves shorted; pinnae and interpinnae modified.
ala--albina ala1, alb1White or cream-colored cotyledons.
ala-2--albina-2 alb2, ala2White or cream-colored cotyledons.
ala-3--albina-3 ala3, alb3White or cream-colored cotyledons.
ala-4--albina-4 ala4, alb4White or cream-colored cotyledons; lethal.
alb--albescent  White variegation promoted by low temperatures.
albprov2albescentprovisional 2albSame as alb.
alc--alcobaca  Greatly delayed fruit ripening processes.
alu--alutacea alu1Small plant, yellowish foliage.
alv--alvoviridis alb, alb1Whitish green cotyledons, pinnae; strong anthocyanin in stems, veins.
Am--Alfalfa mosaic  Resistance to alfalfa mosaic virus
ama--albomarginata  Shorter internodes; leaves small and spotted with yellow-green or yellow; flowers small and petals narrow with white edges.
an--anantha an^1, an^2, caInflorescence exceptionally large and excessively branched; no flowers appear, instead the axes of the inflorescences ramify into smaller and smaller segments, continuing such growth throughout the season; flowers reduced to small vestigial structures; heterozygotes are somewhat intermediate.
an-2--anantha-2  Inflorescence highly branched, terminates in very compact ovary-like masses of tissue; aborted flowers.
an-3--anantha-3  Leaves small, light green; inflorescence indefinitely ramifying, no normal flowers, resembling cauliflower; completely sterile.
ana--angustata ana1Shortened leaves, stems; pinnae small, narrow, pointed; smaller flowers, narrow petals.
ang-2--angustifolia-2 ang2Smaller, compact, slow-growing plant; narrow, light-green pinnae.
anr--antirrhinum  Fruit elongated with hollow tip from which epiglotis extends resembling a snapdragon flower.
ant--aurantia ant1Short thick stems; many short side shoots; narrow, rugose, light green pinnae; light orange fruit; colorless pericarp.
anu--angusta anu1Dark gray-green pinnae narrowed to 1/3 normal width; short leaves.
ap--apetalous  Corolla reduced or absent; pollen scarce and nonfunctional
apl--applanata  Broad, spreading habit; long, laxly pinnate leaves; heterozygote intermediate.
apn--albo-punctata  Fine white speckling uniformly on cotelydons; mature leaves small, wilty; tiny flowers.
Aps-11Acid phosphatase-1   
Aps-12Acid phosphatase-1   
Aps-1nAcid phosphatase-1   
Aps-21Acid phosphatase-2   
Aps-22Acid phosphatase-2   
Aps-23Acid phosphatase-2   
Aps-2nAcid phosphatase-2   
ar--arrecta  Small, compact plants; leaves yellow-green, virescent.
are--anthocyanin reduced  Almost no anthocyanin until fruits set, then moderate amount in young foliage.
ari--aridifolia  Plants 1/3 normal size; cotyledons small, pointed, dull dark grey-green; die prematurely.
as--asynaptic as1Asynaptic meiosis; high pollen and ovule sterility.
as-2--asynaptic-2 as2Asynaptic meiosis; high pollen and ovule sterility.
as-3--asynaptic-3 as3Asynaptic meiosis; high pollen and ovule sterility.
as-4--asynaptic-4 as4Asynaptic meiosis; high pollen and ovule sterility.
as-5--asynaptic-5 as5asynaptic meiosis; high pollen and ovule sterility.
as-6--asynaptic-6 as6Delayed and asynaptic meiosis; chromatin degeneration; possibly two loci involve
Asc--Alternaria stem canker resistance  Resistance to Alternaria stem canker
at--apricot  Flesh of ripe fruit yellow with pinkish blush; carotenoid content similar to r/r.; corolla pale yellow, nearly white.
atn--attenuata atNarrow, pointed leaves, yellow-green, purple below; rigid cylindrical habit; thin shoots.
atv--atroviolacium  Excess anthocyanin on leaves, stems, and fruits.
au--aurea  Bright yellow cotyledons and leaves; seedling etiolates; plant vigorous.
au(1k)aureaKnowles  
au(1s)aureaStubbe'sau^2, au, bracBright yellow cotyledons and leaves; some times whitish; corolla pale yellow; moderately vigorous in condine red; seedling etiolates; plant vigorous.
au(2k)aureaKnowles second  
au2aurea au^3Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au3aurea au-2, au2Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au6aurea yg^6, yg-6, au^yg-6, yoBright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
autlaureaTorrey Lyons' Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
auwaurea w616Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
auc--aucta  Few branches; leaves broad, rounded, later becoming blistered, dark green; number of flowers and fruits reduced.
aur--aurantiaca aur1Small, pointed, yellowish light-green pinnae; orange fruit.
aus--austera  Plants always 1/3 to 2/3 normal size; pinnae finely divided, in some cases bent upwards; foliage darker and duller than normal.
aut--aureata  Vireescent, flush of gold near growing point in early development, not detectable in older plants.
auv--aureate virescent  Seedling 'gold dust' virescence; leaf segments pointed, sometimes fimbriate. Mature plant nearly normal.
avi--albovirens avi1Smaller plant; foliage variously variegated yellow, white, grey-green; reduced fruit set.
aw--without anthocyanin aba, ab, a179Free of anthocyanin, same phenotype as a.
aw(1k)without anthocyanin aw^2Completely free of anthocynanin. Radiation induced in Money Maker.
awprov3without anthocyaninprovisional 3awFree of anthocyanin, same phenotype as a.
awprov4without anthocyaninprovisional 4awFree of anthocyanin, same phenotype as a.
awprov5without anthocyaninprovisional 5awFree of anthocyanin, same phenotype as a.
aw-2--without anthocyanin-2  No anthocyanin at any stages; reduced growth rate, dense bushy habit.
B--Beta-carotene  Flesh of ripe fruit orange, due to high B-carotene, low lycopene concentrations.
BcBeta-carotenecrimsonog^c,Crn,Cr,crn-2,cr-2Increased fruit lycopene content; phenotype similar to B^og.
BmBeta-caroteneminutum High B-carotene, low lycopene in ripe fruit. .
BogBeta-caroteneold goldogCorolla tawny orange; increased fruit lycopene.
bc--bicolor biYoung leaves mottled brownish olive normal, later glazed dark green; short internodes; heterozygote intermediate for some characters.
Bco--Brilliant corolla  Corolla pigment intensifier.
be--blue-green  Small plant with blue-green leaves.
bg--bursting resistance  Resistance of fruit to bursting.
bi--bifurcate inflorescence  With f and j causes extreme fasciation of the "ring type".
bip--bipinnata  Leaves highly divided, with smaller, acute segments.
bipprov2bipinnataprovisional 2bipLeaves highly divided, with smaller, acute segments.
bip-2--bipinnata-2 bip2Slender, pointed, light gray-green cotyledons; doubly pinnate, rugose leaves.
bk--beaked  Fruits with sharp points at stylar end.
bk2beaked  Fruits with sharp points at stylar end.
bk3beaked  Fruits with sharp points at stylar end.
Bk-2--Beaked-2  Sharp point at stylar end of fruit
bks--black seed bks1-1Black seed testa, reduced germination.
bks2black seed bks1-2Black seed testa, reduced germination.
bl--blind  Growth of axis terminates in single flower or inflorescence as if an extremem expression of sp (self-pruning); no buds in axils of leaves or cotyledons; late season adventitious shoots may emerge from the midribs; expression stronger in field than greenhouse.
bl2blind to^2Determinant growth; few flowered inflorescences; fasciated flowers; enlarged, leafy calxy; less extreme than bl.
bltoblindtorosatoDeterminate habit; firm, strongly uprolled pinnae; fasciated flowers and fruits; expression stronger in field.
bls--baby lea syndrome almAnthocyaninless; restricted root system; short internodes, leaves, and trusses
blsprov2baby lea syndromeprovisional 2blsAnthocyaninless; restricted root system; short internodes, leaves, and trusses
blsprov3baby lea syndromeprovisional 3blsAnthocyaninless; restricted root system; short internodes, leaves, and trusses
bn--blunt  Stamens absent, rarely present, with functional pollen, in greenhouse. Usually petaloid stamens adnate to pistil; leaf segments obtuse
bnstsbluntStamen suppressorsts, sl-6Stamens absent, rarely present, with functional pollen, in greenhouse; fewer blunt leaflets; with sp:+ 6-10 leafs between inflorescences.
Bnag-11Beta-N-acetyl-D-glucosaminidase-1   
br--brachytic  Internodes shortenedm resulting in low stature and compact growth habit; by the third-leaf stage, br plants are only half as tall as +; differs from dwarf (d) in having normal-colored, nonrugose foliage, and normal instead of sessile and compact leaf segments.
brt--bushy root  Radical branches early; radical and root profusely branched; root tips twisting upwards, not dense growth.
brt-2--bushy root-2  severely stunted growth (1/20) dense bushy growth of twisted roots.
bs--brown seed  Brown seed color behaving as endosperm trait.
bs-2--brown seed-2  Brown seed color governed by genotype of endosperm.
bs-3--brown seed-3  Brown seed color determined by genotype of endosperm.
bs-4--brown seed-4  Brown seed color controlled by genotype of endosperm.
Bt--bursting digenic inheritance bgSusceptibility to fruit bursting.
btl--brittle stem  Growth ceases with first flowering; stems brittle, with progressive defoliation; temperature-sensitive and expressed only under field conditions; B inefficient.
bu--bushy fruHypocotyl normal, but cotyledons cuneate and held close to the plumule; first three internodes very short, resulting in dwarfing of the seedling to half height of normal; in later stages, internodes shortened, with excessive production of side branches, giving a bushy appearance; all parts of inflorescence foreshortened; abscission joint of pedicel imperfect and shifted towards calyx.
buabbushyabbreviatafru^abInternodes and inflorescences shortened; long petioles.
bucinbushycompact inflorescencecinInternodes and inflorescences shortened; long petioles.
bucin-2bushycompact inflorescence 2cin-2Internodes and inflorescences shortened; long petioles. Duplicate genes determining character similar to bu.
buhembushyhemiglobosafru^hemInternodes and inflorescences shortened; long petioles. Phenotype like bu except leaves longer, more lax; heterozygote intermediate in some traits.
bul--bullata  Leaf surface blistered and chlorotic with a network of darker veins; short internodes.
buo--bullosa buo1Plants and seedlings larger in all parts; broad and rugose pinnae.
Bw-1--Bacterial wilt-1   
Bw-2--Bacterial wilt-2   
Bw-4--Bacterial wilt-4   
Bw-5--Bacterial wilt-5   
c--potato leaf  Number of leaf segments reduced
cintpotato leafintegerrimaintNumber of leaf segments reduced
cprov2potato leafprovisional 2cNumber of leaf segments reduced
cprov3potato leafprovisional 3cNumber of leaf segments reduced
cprov4potato leafprovisional 4cNumber of leaf segments reduced
cprov5potato leafprovisional 5cNumber of leaf segments reduced
cprov6potato leafprovisional 6cNumber of leaf segments reduced
car--carinata  Shortened, keeled pinnae, dark gray-green color; deformed flowers; elongated fruit.
car-2--carinata-2 car2Smaller plant; slender stems; smaller pinnae, nearly entire margin.
cb--cabbage  Compact habit; leaves large and dark green; fewer locules.
cb-2--cabbage leaf-2  Larger plant parts; leaves epinastic and blistered; broad, rugose leaves, similar to dwarf.
ccf--cactiflora  Flower greatly fasciated; and roecium gynoecium and green tissues fused and distorted, enlarging into fruitlike structure; complete make and female sterility.
cd--condensata  Seedling small; internodes shortened; leaf light grey-green with narrow, keeled segments; fruit round and relatively large.
ce--cernua ce1Cotyledons lightly dotted in whitish yellow-green; lax, inclined plant.
Cf-?--Cladosporium fulvum resistance  Resistance to specific races of Cladosporium fulvum
Cf-1--Cladosporium fulvum resistance-1 Cf, Cf1, CfscResistance to race 1 and 3 of Cladosporium fulvum.
Cf-13Cladosporium fulvum resistance-1 Cf-5, Cf5Resistance to specific races of Cladosporium fulvum.
Cf-2--Cladosporium fulvum resistance-2 Cf2, Cfp1Resistance to races of Cladosporium fulvum.
Cf-3--Cladosporium fulvum resistance-3 Cf3, Cfp2Resistance to specific races of Cladosporium fulvum
Cf-4--Cladosporium fulvum resistance-4 Cf-8, Cf4, Cf-1^2Resistance to specific races of Cladosporium fulvum.
Cf-6--Cladosporium fulvum resistance-6  Resistance to specific races of Cladosporium fulvum.
Cf-7--Cladosporium fulvum resistance-7  Resistance to specific races of Cladosporium fulvum.
Cf-9--Cladosporium fulvum resistance-9  Resistance to specific races of Cladosporium fulvum.
cfa--conferta cfa1Very small, compact bush; strongly shortened internodes, young leaves.
cg--congesta cg1Small, dense bush; late growth retardation; short internodes; pointed primary leaves.
ch--chartreuse  Greenish-yellow subnormal petals; reduced fertility.
chln--chloronerva  Upper leaves smaller and chlorotic interveinally; strong graft response.
chrs--chrysanthemum sterile  Flowers fascicled 6cm across; many small pale yellow petals; stamens dialytic and without pollen; fruits deformed, orange-red fruits with brown sectors; poor fertility.
ci--cincta ci1Small, irregular bush; many side shoots of equal length from lower leaf axils.
cir--circumsaepta cir1Rounded, light gray-green pinnae, downrolled margin.
cit--citriformis  In pm stock; Plant reaches 1/3 - 2/5 typical size; half erect, weakly branched; foliage color grey-green; terminal segment of primary leaves long and narrow; older leaves serrated; lemon shaped fruit with conspicuous stylar scar.
cjf--conjunctiflora  Flowers joined in pairs or triplets, otherwise malformed.
ck--corky fruit  Pericarp cracked, exposed tissues becoming brown and leathery.
Cl--Cleistogamous Cl1Flowers fail to open.
cl-2--cleistogamous-2 cl2Flowers open only slightly; reduced fertility.
cla--clara  Yellowish-green leaf color, purple veins and petioles; incompletely dominant.
clau--clausa ff, vcPartly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clauffclausa  Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clauicsclausaincisifoliaicsPinnae margins deeply incised, dentations acute.
clauprov2clausaprovisional 2clauPartly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clauvcclausaveined cotyledon Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits. Original description: cotyledons distinctively veined, leaflets slightly convex and twisted, shallow lobes; growth normal, fertility reduced.
cle--collecta  Reduced growth rate; internodes shortened; small compact bush habit; leaf segments large, rounded, foliolules fewer.
cls--clarescens  Foliage light yellow green early, becoming whitish green and blistered later, brilliant yellow green virescent; plants 1/4 typical size; growth very weak in field, setting fruits only in greenhouse.
clt--coalita  Size and habit typically spl; primary leaves with entire end segments and only tiny laterals; mature leaves typically 5-segmented through fusion with end segment; segments always entire; resembles but is not allelic to sf.
clt-2--coalita-2  Terminal pinnae large, second order pinnae few and masked by 1st order pinnae, little serration.
cm--curly mottled  Strong virus-like mottling and distortion of leaves and abortion of flowers. Highly sensitive to environment, particularly temperature; highest expression and nearly complete sterility under low light, cool conditions of winter, whereas phenotype essentially normal in mid-summer.
cma--commutata  dwarf with bright green foliage
Cmm--Clavibacter michiganensis resistance  Resistance to Clavibacter michiganensis.
Cmr--Cucumber mosaic resistance  Resistance to isolate 113A of cucumber mosaic virus, from L. chilense.
cn--cana caCotelydons and leaves gray-green especially on undersides; tiny and unbranched plants.
cna--citrina  Foliage weak grey-green; petals light yellow; anthers greenish yellow.
co--cochlearis  Reduced gray-green leaves with concave spoon-shaped leaflets; small plants of normal habit.
co-2--cochlearis-2 co2Small leaves; deeply crenate, initially gray-green pinnae.
coa--corrotundata coa1Plant small; pinnae fewer, broad, rounded, darker; flower parts short and broad.
com--complicata  Smaller, weakly branched erect plant; firm and keeled pinnae.
comincomplicataindigainSmall, dainty, nearly unbranched plants; shortened light gray-green leaves.
com-2--complicata-2 com2Very small plant; short leaves, internodes; few side shoots; purplish shoot tips.
con--convalescens  Yellow-green, paler at growing point; variably depressed growth.
con-2--convalescens-2 con2Narrow cotyledons, dull light gray-green foliage, weakly yellow-green in growth zones.
cor--coriacea  Smaller, unbranched plant; all parts reduced; short leaves with plicate segments.
cp--compacta cp1Short leaves, smaller, dark green pinnae.
cpa--composita cpa1Large, strongly branched, leafy inflorescence; jointless pedicel.
cpf--cupuliformis  Cotyledons fused, cup-shaped, becoming coarse and enlarged later; transition to mature leaves without primary leaves; plants die at 3-4 leaf stage; line maintained via heterozygotes.
cpf-2--cupuliformis-2  Phenotype similar to cpf.
cpf-3--cupuliformis-3  Cotyledons adnate to form a cup; no primary leaves develop; maintained via heterozygotes.
cps--compactilis  Plant smaller at all stages; leaves somewhat wider, lightly blistered and darker green.
cpt--compact  Highly branched, compact plant; lax stems; pale, smaller leaves; reduced fruit set.
cr--radial crack resistance  Resistance of fruit to radial cracking.
Cri--Crispa  Very small in all parts; poorly viable without grafting; leaves light olive green, necrotic. Cri/+ necrotic spots on pinnae centers.
Crk--Crinkled  Leaves like those of Cu but less condensed and segments dentate; homozygote lethal.
crt--cottony-root  Overgrown root hairs with cottony appearance
cru--corrupta cru1Smaller plant, irregular, sometimes distorted growth; spreading necrosis; firm, leathery, involuted brittle pinnae.
cry1--cryptochrome 1  Hypocotls and stems are elongated, and anthocyanins reduced; seedlings are de-etiolated under blue light; phenotype more extreme in combination with phyA and phyB1.
cs-2--corollaless-2  Apetalous; dialytic anthers; exserted stigma; self fertile, rarely sets fruits naturally.
cta--contaminata cta1Smaller, flat, broad bush; many side shoots; at end of season stems and leaf midribs turn necrotic and brittle; also reduced fertility.
cta-2--contaminata-2 cta2Leaves 1/4 normal size; stems with corky surface, variable necrosis of vascular tissue.
ctb--cotyledon tops bifurcation  Bifurcation of cotyledon tops by a central vein.
ctr--citrate concentration  Low citric acid in fruit (cv. Tondo Liscio) vs. high concentration, ctr (PI 263713 = L. esculentum).
ctr1citrate concentration   
ctt--contracta  All shoots somewhat shortened, resulting in round bush habit; pinnae weakly serrated, plicate, and flattened; foliage somewhat lighter, and growing points are somewhat darker.
ctt-2--contracta-2  Plant half normal size at flowering; internodes shortened; leaves are narrow, pointed and blistered.
Cu--Curl  Mid- and lateral veins and petiole greatly foreshortened producing crumpled leaf; homozygous viable.
cu-2--curl-2 cu2Small, downcurled leaves; resembling Cu.
cu-3--curl-3  Dwarf, short and thick hypocotyls, curled cotyledons, and dense curly leaves; reduced fertility.
cu-3abscurl-3altered brassinolide sens. Similar to cu-3, no restoration by brassinolide application
cua--curta  Seedling and mature plant smaller, round bush habit; internodes and leaves shortened; leaves rounded and wider.
cul--culcitula  Seedlings dark green with strong anthocyanin, later foliage dark grey-green; plants reach 1/3 to 2/5 typical size; internodes very short, habit cushion-like.
cup--cupida cusExtremely susceptible to drought damage at all stages; leaves small, yellowish, wilting, and eventually necrotic.
cur--curvifolia  Leaves twisted, yellow-green; short internodes.
cv--curvata cuYoung stem curves from node to node; dark rugose leaves; dwarf compact habit; heterozygote intermediate for some traits.
cv2curvata acuSmall plant; leaves, pinnae briefly stalked; leaf surface furrowed; older leaves strongly bent downward.
cv-2--curvata-2 cv2Twisted cotyledons; short, rounded leaves; rugose, yellow-green pinnae.
cva--conversa  Small, dense plant; variable fading of foliage.
cva-2--conversa-2 cva2Small, compact plant, leaves, pinnae.
cvl--convoluta cvl1Small, dense plant; short internodes; pinnae of oldest leaves involuted.
Cvx--Convexa  Seedling and mature plant with narrow convex leaves, deeply veined and tending to roll downward on margins.
cy--cyathiformis  Cotyledons united at one or both edges; lateral shoots suppressed; pinnae margins nearly entire.
d--dwarf  Shortened hypocotyl, darker, broader and shorter cotyledons; stems heavy and erect; plant compact, internodes shortened to about 2.5cm; leaves very distinct, with reduced number and size of segments; dark green color, puckered rugose surface, down-curled magins, and broader, shorter outline of whole as well as individual segments; similar reduction in size and foreshortening of inflorescence, flowers and fruit.
dbdwarfbroccoli Extreme dwarf; more stunted than d^x; leaves reduced to small, dark green globs of curled, rugose tissue; flowers reduced, with tiny corolla segments.
dcrdwarfcrispatarob^crispAllele of phenotype intermediate between d and d^x.
dimdwarf rob^immAll parts foreshortened; leaves dark and rugose.
dprov2dwarfprovisional 2dAll parts foreshortened; leaves dark and rugose.
dprovcr-2dwarfcrispatad^crAll parts foreshortened; leaves dark and rugose.
dprovcr-3dwarfcrispatad^crAll parts foreshortened; leaves dark and rugose.
dprovx-2dwarfextreme-2d^xAll parts foreshortened; leaves dark and rugose.
dxdwarfextreme Resembles dwarf (d) but more extreme in all respects; growth is very slow; all parts of plant reduced in size and most organs modified in the direction of shorter and broader shape; leaves very dark green, the surface coarsely bullate and midrid twisted; plants are moderately fertile; recessive to d and to +.
d-2--dwarf-2 rob2, rob II, d2Shortened internodes and leaves; thick, rugose, bluish gray-green leaves; not allelic with the mimic,d1.
d-3--dwarf-3 d3Smaller, sturdy, erect plant; all parts foreshortened; rugose and dark-green leaves.
d-4--dwarf-4 d4Mildly dwarf type; broad, shortened, crowded, darker pinnae; darker flower color.
d-5--dwarf-5 d5, d2, M-5Slow growing plant with normal stem and leaf proportions; semi-sterile; classification good when three weeks old.
d-6--dwarf-6 d6, d5Compact and small plant; rugose, and keeled pinnae, upper surface shiny dark green.
d-7--dwarf-7  Plants reduced to 10-12 inches tall; short internodes; leaves highly divided with strong anthocyanin.
d-8--dwarf-8  Plants reduced to 6-8 inches tall, short internodes; normal flowers but no fruit set in field.
dc--decomposita dc1Slender, strongly sinuous and crenate pinnae; more secondary pinnae; micromutation.
dd--double dwarf d^xxExtremely retarded and highly modified dwarf.
de--declinata  Great reduction in size of all parts; leaves dark yellow-green, leaflets roll toward underside.
deb--debilis  Leaves emerge yellowish; necrotic mottling, whitish, and later brown, often constricting leaves in middle.
dec--decumbens  Lax and decumbent habit; early fruiting.
dec-2--decumbens-2 dec2After start of blooming plant becomes decumbent.
def--deformis  Seedlings normal, later leaves becoming progressively reduced, filiform, and with adnate segments; deformed flowers and reduced fertility.
def2deformis vitLeaves become progressively more deformed with twisted filiform leaflets.
def-2--deformis  Like def; leaves deformed after flowering.
Del--Delta  Reddish-orange mature fruit color, due to inhibition of lycopene, and increase of delta-carotene.
deli--deliquescens  Early habit dense later loosely parting; shortened yellowish leaves with narrow acute segments.
deli-2--deliquescens-2 deli2Smaller compact bush; short internodes and leaves; all foliage is light yellow-green.
den--densa  Typical spl seedlings; plant 3/4 normal size; stems and petioles strong; internodes somewhat shortened; leaf segments large and coarse; foliolules blistered; foliage color somewhat lighter.
dep--deprimata  Slightly depressed growth; dainty, darker leaves.
depa--depauperata  Very small plants with short internodes; leaflets bowed, color variable, and mostly yellowish.
det--detrimentosa  Small, weakly branched and lax plant; yellowish foliage.
det2detrimentosa  Small, plant; dainty, pale yellow-green leaves; slender pinnae; less discoloration and premature dying than det.
det-2--detrimentosa-2  Plants reach 1/4-1/3 normal size; cotelydons narrow and tapered; leaves are shortened, segments small, keeled, margins serrated, and dull dark grey-green.
Df--Defoliator  Homozygous seedlings retarded in growth, dark green, and die early; heterozygous are viable, but leaves become necrotic and fall prematurely.
dfd--delayed fruit deterioration  Fruit softening greatly reduced; shelf life up to seven months; resistant to postharvest disease
dgt--diageotropica lz-3Plant habit prostrate due to reduced gravitropic response; growth retarded; stems and leaves droopy; cotelydons concave. Roots grow horizontally rather than downwards.
dgtdpdiageotropicadrooping leafdpLeaves drooping, elongate, dark green; stems weak, slender, and prostrate.
Dia-21Diaphorase-2   
Dia-22Diaphorase-2   
Dia-31Diaphorase-3   
Dia-32Diaphorase-3   
Dia-41Diaphorase-4   
dil--diluta  Leaves dull light green, small, and roughened; plants half normal size.
dil-2--diluta-2 dil2Light colored, denser plant; short leaves, smaller pinnae.
dim--diminuta  Somewhat smaller plants and leaves, shortened internodes; older leaves gray green with violet veins.
dim-2--diminuta-2 dim2Very small plant, leaves, pinnae; much anthocyanin in hypocotyl, growth zones.
dis--discolor  Leaf color light green with virescent growing regions, also with darker veins.
dis-2--discolor-2 dis2Cotyledons first yellow-green, then whitish light-green; small compact plant.
div--divaricata  Small squarrose plant with intercostally yellowish leaves and ventrally purple.
dl--dialytic  Large trichomes partially suppressed and distorted; stamens not united, resulting in partially reduced fruit set from self pollination; seedlings can be distinguished in the 2- or 3-leaf stage by the shortening, bending and forking of the long and intermediate hairs.
dlSdialyticSolanumDl^sAnthers free, but trichome modifications of original dl absent. Expression is dominant to wild type.
dlb--dilabens dlb1Small, compact plant with shortened yellowish leaves.
dlu--dilucida  Plants reach 1/8-1/6 normal size, dwarf bush habit; foliage bright light green, light yellow-green virescent; petioles, stems, veins light violet; very low fruit set.
dm--dwarf modifier d2Causes extreme dwarfing.
dmd--dimidiata  Hypocotyl is strong; cotelydons are almost oval and arched downwards; internodes short and result in is small plants; leaf segments broad, coarse, and flat with 1 - 2 pairs of laterals grey-green; few foliolules develop later; corolla segments broad and flat; fruits few and seedless; maintained via heterozygotes.
dmt--diminutiva  Shortened plant height and length of stem internodes, leaves, and pinnae.
dpa--deparca  Small seedling and mature plant; little branching; leaf lighter at base, smaller, dark green; strong anthocyanin.
dpi--densipinnata  Plant 1/2-3/4 normal size; small bush habit; cotyledons narrow with tapered tip; leaves dull grey-green; leaf segments large, crowded and fewer foliolules.
dpr--depravata  Cotelydons and primary leaves strongly curled; internodes and leaves shortened; young leaves very asymmetrical, later deformed with long petiole.
dps--diospyros  Fruit tissue is dusky orange.
dpy--dumpy  Leaves like those of d:x but internodes longer; Leaves greatly condensed, rugose, dark green; internodes somewhat foreshortened
dpyprov2dumpyprovisional 2dpyLeaves like those of d:x but internodes longer.
dpyprov3dumpyprovisional 3dpyLeaves like those of d:x but internodes longer.
drt--dwarf root  Reduced root system
ds--dwarf sterile  Stunted plants with short internodes and reduced leaves; partially male-sterile.
dt--dilatata dt1Plant small; pinnae yellowish; veins darker with uprolled margins.
dtt--detorta  Cotyledons epinastic, irregularly crooked, many seedlings with 3 cotyledons; plants 2/5-3/5 normal size, less erect than normal; stems crooked and leaves and pinnae disordered; pinnae partly plicate, partly spiraled.
du--dupla  Young plants typically splendens, later reach 2/3 size; habit half erect; branching reduced, particularly in the side shoot below the first inflorescence; leaf segments broad and doubled in number with many foliolules; strong anthocyanin in shoots exposed
dv--dwarf virescent  Pale green virescent; plants always stunted.
dx-2--extreme dwarf-2  Resembles d:x in most respects; all parts extremely shortened; plant erect with no branching, attains half normal size; leaves very rugose and bright dark green above; flowers are small.
e--entire bFirst true leaf undivided but serrate, mature leaves usually have reduced number of lateral segments, which have variably serrate margins; contraction indicated by crowding of lateral segments and distortion of midvein.
e2entire  Fewer fused leaf segments and distorted midvein, entire or broad leaflets
eprov3entireprovisional 3eFewer leaf segments; midvein distorted
e-2--entire-2  e-type leaves
ec--exserted carpels  Fruits vary from none to many exserted carpels.
eca--echinata  Early growth typically pm and later reaching 2/3-3/4 typical size; from fruit set on main stem and branches grow vertically with dense to loose bush habit, lending porcupine-like appearance.
el--elongated eElongated fruits as in Oxheart.
ela--elata  Plants reach 3/4 normal size; dense, erect, and bush habit; internodes shortened; leaf segments large, wide, and strongly blistered.
ele--elegans  Leaves reduced, yellow-green and dainty; plants tiny
ele2elegans angReduced, sometimes chlorotic, narrow leaves; slow growth.
ele-2--elegans-2 ele2Narrow, rounded cotyledons, margin rolled up; smaller, light yellow-green pinnae.
elo--elongata elo1Elongated, weakly branched stems, initially erect, become procumbent.
elu--eluta  Yellow green leaves; strong branching tendency; slight growth depression.
elu-2--eluta-2 elu2Yellow-green to light gray-green leaves; late in season strong anthocyanin in shoots, stems, and veins.
em--emortua em1Small plant; reduced side branching; spreading necrosis of older leaves leading to their premature death; expression best in field.
en--ensiform  Sword-shaped sepals.
ent--entire leaflet  Leaflets with entire margin.
ep--easy peeling  Epidermis can be peeled from fruit without pretreatment.
Epi--Epinastic  Severe epinasty of leaves, swelling of stems and petioles, prolific branching of roots. Partially dominant.
er--erecta  Dwarf bushy plants with short internodes; shortened dark green leaves; heterozygotes intermediate.
er12.1--earliness   
er-2--erecta-2  Plant vigorous, branches erect, internodes slightly shortened; leaves shorter, segments smaller, keeled and blue-green at tips; many flowered.
era--eramosa era1Smaller, weakly branched plant; whitish zones in primary and later in leaves causing irregular pinnae development.
Erl--Erectoid leaf  Leaves held at acute angle on stem; partially dominant.
Est-11Esterase-1   
Est-12Esterase-1   
Est-13Esterase-1   
Est-14Esterase-1   
Est-15Esterase-1   
Est-1nEsterase-1   
Est-21Esterase-2   
Est-31Esterase-3   
Est-41Esterase-4   
Est-42Esterase-4   
Est-43Esterase-4   
Est-44Esterase-4   
Est-45Esterase-4   
Est-46Esterase-4   
Est-47Esterase-4   
Est-48Esterase-4   
Est-51Esterase-5   
Est-61Esterase-6   
Est-71Esterase-7   
Est-72Esterase-7   
Est-81Esterase-8   
ete--extenuata ete1Slender shoots; slender and thick pinnae; irregular growth; poor seed set.
eu--eugenol concentration  High eugenol concentration in fruit.
ex--exserted stigma  Exserted stigmas, or styles are twisted within anther tubes.
exa--expassa exa1Branches partially decumbent; yellowish gray green older leaves turning yellow prematurely.
exl--exilis exVery small plant; tiny plicate leaves of light gray-green color.
exs--excedens exs1Smaller bush; short internodes, young leaves; older leaves leathery, and shiny.
f--fasciated fruit  Increased number of locules, stamens, and segments of the corolla and calyx; ovary and fruit more flattened and irregular in outline.
fDfasciated fruitDominant Fruits many-loculed; phenotype more extreme than original allele of f; carpels unfused; dominant allele.
fa--falsiflora fa1Giant, vegetative, and highly ramified inflorescence; completely sterile.
far--farinosa far1Smaller, weakly branched plant; many, deeply crenate, wavy, gray to yellow-green pinnae.
fc--flaccescens  Internodes and leaves shortened; cotyledons narrow, wavy, light green; plants retarded at flowering, reach 1/5 normal size; leaf segments small, dark grey-green, tending to shrivel and die early.
fcf--fucatifolia fcf1Small spreading plant; yellow or light green cotyledons, primary leaves, and lower leaves at later stages.
fd--flecked dwarf  Retarded at all stages; leaves flecked with light green.
Fdh-11Formate dehydrogenase-1   
Fdh-12Formate dehydrogenase-1   
fe--fertilis  Leaves roll dorsally; fasciated flowers, fruits.
fer--fe inefficient  Severe chlorosis beginning in first true leaves due to faulty iron transport in xylem.
FgrHFructose to glucose ratiohirsutum Ratio of fructose to glucose in fruit approx. 1.5:1; partially dominant over L. esculentum allele
fgv--fimbriate gold virescent  Leaf fimbriate margined, ephermerally "gold dust" virescent.
fi-2--filiforme-2  Phenotype like Schiemann's (1932), filiforme, equivalent to extreme wiry phenotype; cotyledons extremely narrow, grey-green; primary leaves also very narrow, later some leaves have only midveins, others with modified terminal segments and some epetiolate lateral segments; corolla with filiform segments; internodes very short, plant height averaging 20 cm.; no fruit set, line being maintained via heterozygotes.
fil--filaris  Habit strongly branched and bushy; cotyledons slightly wavy; leaves wide, rounded, and often asymmetric with elongated petiole and filiform terminal segments; inflorescence very compound, petioles narrow, and fruit elongate.
fir--firma  Broad blunt leaf segments; bushy cylindrical plants; foreshortened inflorescence.
fir-2--firma-2 fir2Sturdy, compact seedlings; broad, rounded, rugose and dark green pinnae; small dark flowers.
fir-3--firma-3 fir3Very small, compact bush; small, dark green, and rugose pinnae.
fis--fistulosiflora  Corolla does not open although segments not fused.
fl--fleshy calyx  Sepals fleshy, often curled.
fla--flavescens  Leaves light with few segments; variably reduced plant size.
flav--flavida  Leaves yellowish, mottled yellowish green, and small; heterozygote intermediate for some traits.
flav-2--flavida-2 flav2Smaller, compact bush; slender stems, many side shoots; leaves yellowish light green.
flc--flacca  Leaves overwilt under dry or sunny conditions, leading to necrotic areas; plant small, erect, nearly unbranched, and later spreading.
fld--flaccida fld1Small, weakly branched and erect plant; darker and involuted wilty pinnae, dying prematurely.
fle--flexifolia fle1Very small plant, leaves with strong anthocyanin, borne at acute angle to stem in young plant.
fli--filiform inflorescence  Filiform outgrowths and cupped leaves in late or terminal inflorescences; narrow corolla.
fms--female-sterile  Distorted, poorly developed, highly sterile gynoecium; normal pollen production.
fn--finely-netted  All true leaves light green with slightly darker veins.
fp--fruit pox  Many small dark green spots on immature fruit, rupture prior to ripening.
fr--frugalis  Small compact bush; older leaves dark-green; brittle, shiny.
fra--fragosa fra1Smaller, very brittle bush; broad cotyledonsc pinnae; heterozygote recognizable.
frg--fragilis frg1Smaller, erect, nearly unbranched plant; short, dull light green leaves, yellowing prematurely.
fri1far red light insensitive phyAInsensitive to far red light; hypocotyls elongated relative to wild type under far red light.
Frl--FORL resistance Fr1, Fr-1Resistance to Fusarium oxysporum f. sp. radicis- lycopersici; incompletely dominant.
Frly--Frilly FrlLeaf with serrate undulate margins.
fro--frondea  Seedlings and plant habit and size normal; inflorescence highly proliferated; pedicels jointless; 4-parted corollae; segregation deficient in mutant homozygotes.
Frs--Frosty spot NecHypertrophic pustules along leaf veins followed by chlorosis then necrosis.
frt--fracta  Plants attain 1/4 normal size; habit upright, weakly branched; leaf margins later turned upward; foliage dark green, shoots brittle.
Fs--Fruit stripe  Fruit with dark green radial stripes opposite locules
fsc--fuscatinervis dkvLeaves yellow-green; veins always darker green.
ft--fruiting temperature  Fruits can set at 40F.
fu--fusiformis  Early growth habit is bushy and spindle shaped; leaves shortened, curved upwards, shiny yellowish, and paler at growing point.
fua--fucata fua1Smaller plant; young pinnae yellow green, darker veins; older foliage variably lighter colored.
fug--fulgida fug1Small plant; foliage variably whitish or yellow-green, sometimes zonally marked.
ful--fulgens  Bright yellow virescent, older leaves becoming greener; golden yellow in field; yellow unripe fruit color.
ful2fulgens ful1^2Foliage at all stages light-green to light yellow-green; stems and leaf mid-ribs sometimes purplish.
ful-2--fulgens-2 ful2Small plant; short internodes; light-green foliage, yellow-green growth zones.
ful-3--fulgens-3  Stunted virescentseedlings, overall brilliant yellow; closely resembling ful; mature plant bright yellow green.
fus--fulgescens  Seedlings normal; plants normal size; foliage light green; growing points lighter; resembles, but is not allelic to ful.
fv--flaveola  Seedlings small; leaf color light grey-green, a flowering light yellow green virescent; older leaves yellowish.
Fw--Furrowed  Deeply furrowed, dark green cotyledons; severely stunted plant; homozygous inviable.
fx--flexa  Young plants typically normal; main stem bends at start but becomes erect later, lending the plant a prostrate habit, similar to reptans(re).
fy--field yellow  Foliage yellow at all stages and under field conditions.
g--grooved  Inhibits modifiers, permitting expression of r.
ga--galbina ga1Foliage at all stages variably white-gray-yellow-normal green; environmentally sensitive.
ga-2--galbina-2 ga2Pinnae dark to light gray-green, yellow base; stems at all stages with strong anthocyanin; plant smaller in some years.
gas--gamosepala gas1Small plant; gray-to-light-green foliage; wavy, irregularly crenate and rugose pinnae; partly connate sepals.
gbl--globula  Plants always greatly reduced; hypoctyls; extremely short internodes; strongly branched; pinnae grey-green, coarse, crooked, downward; only observed in greenhouse; no flowers developed; maintained via heterozygotes.
Gdf--Gold fleck  Small dark green spots on immature fruit, do not rupture but turn yellow on ripe fruit.
GecGamete eliminatorCondine Red Most gametes abort when interacting with Ge:p.
GenGamete eliminatorneutral neutral allele found in most genotypes
GepGamete eliminatorPearson Most gametes abort while interacting with Ge:p.
gf--green flesh  Persistent chlorophyll giving ripe fruit purplish-brown color.
gfl--globular flower  Corolla tips remain attached, resulting in globular form.
gfo--grandifolia  Habit flatter; cotyledons and primary leaves variable whitish; leaves and segments large and light grey-green.
gh--ghost abIncomplete chlorophyll deficiency, starting green, later breaking to white.
gh-2--ghost-2  Cotelydons yellowish; most leaves variegated.
gi--gibberosa  Seedling relatively coarse; plant size normal; main stem semi-prostrate with reduced branching; leaves. mostly with 2 pairs lateral segments, very few foliolules, segments almost as broad as long, some terminal segments fused with laterals; less hairy stems and petioles.
gib-1--gibberellin deficient-1  Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-2--gibberellin deficient-2  Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-3--gibberellin-deficient-3  Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-3xgibberellin-deficient-3  Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gil--gilva  Yellow cotyledons; highly variable ratios.
gl--glauca  Leaf segments narrow, acute yellow-green; virescent.
gl-2--glauca-2 gl2Short internodes; small, dull, light gray-green leaves.
gla--glaucicolor  Flat dense bush habit, strongly sidebranched; internodes shortened; foliage dull light grey-green; shoot tips with much anthocyanin; flowers small.
glau--glaucescens  Leaves shortened, dull green to yellowish gray-green; small upright habit.
glau-2--glaucescens-2 glau2Smaller plant; leaves dull gray-green; yellow-green growth zones.
glb--globularis  Internodes and leaves shortened; leaves densely pinnate; yellow tinged.
glb-2--globularis-2 glb2Dense, rounded bush; very short internodes; many side shoots; poor fruit set.
glc--glaucophylla  Small plant; leaves light yellow or gray green, shortened.
glc-2--glaucophylla-2 glc2Some pinnae larger, broader; pinnae light-green with lighter flecks.
glc-3--glaucophylla-3 glc3Small plant and leaves of dull light grey-green.
glf--globiformis glf1Very small, irregular bush; all plant parts reduced; many side shoots; multibranched inflorescences; young heterozygotes recognizable.
glg--galapagos light green  Leaves pale gray-green, darker veins; plant small
glm--glomerata  Seedlings normal; young plant with stronger anthocyanin; plants 1/3-2/3 normal size; shoots become necrotic at fruiting, resulting in condensed habit; later some branches become normal.
glo--globosa  Internodes short; leaves short, pale green; incompletely dominant
glo2globosa inx, introLeaves tend to roll toward upper side, dull gray-light green; shortened internodes; excessively branched.
glo-2--globosa-2 glo2Very compact, rounded bush; many side shoots; short internodes; small flowers, fewer clusters.
glo-3--globosa-3 glo3Full sized erect bush; half sized, slender pinnae.
glu--glutinosa glu1Dark green, shiny fruit with sticky epidermis; poor germination.
gm--gamosepalous  Calyx and corolla segments tending to be connivent.
Got-11Glutamate oxaloacetate transaminase-1   
Got-12Glutamate oxaloacetate transaminase-1   
Got-21Glutamate oxaloacetate transaminase-2   
Got-22Glutamate oxaloacetate transaminase-2   
Got-23Glutamate oxaloacetate transaminase-2   
Got-24Glutamate oxaloacetate transaminase-2   
Got-2nGlutamate oxaloacetate transaminase-2   
Got-31Glutamate oxaloacetate transaminase-3   
Got-32Glutamate oxaloacetate transaminase-3   
Got-33Glutamate oxaloacetate transaminase-3   
Got-3nGlutamate oxaloacetate transaminase-3   
Got-41Glutamate oxaloacetate transaminase-4   
Got-42Glutamate oxaloacetate transaminase-4   
Got-4nGlutamate oxaloacetate transaminase-4   
Gp--Gamete promoter  Strongly increased fertilization of gametes carrying it.
gq--grotesque  Irregular androecium; pistil variably twisted, elongated, ridged, or lobed.
Gr--Green ripe grFruit flesh turns slowly to yellowish green, remains firm; resembles gf, except that center of fruit turns red.
GrNr-2Green ripe  Resembles gf, except that center of fruit turns red.
gra--gracilis  Very small, unbranched, delicate plants; much reduced leaves yellowing at margins.
gra-2--gracilis-2 gra2Very small plant, pinnae, short internodes and leaves.
gra-3--gracilis-3 gra3Plant very small, erect, nearly unbranched; short internodes and leaves; pinnae and seedlings smaller.
grc--gracillama grc1Smaller, weakly branched, half-erect plant; yellow-green, slightly involuted pinnae.
grf--grandifructa grf1Broad, lax habit; fewer, large, flattened fruit; heterozygote intermediate.
grl--gracilenta grl1Smaller, erect, weakly branched plant; large pinnae, close together, irregularly yellow green, normal green veins.
grl-2--gracilenta-2  Slender, pointed, pale green cotyledons; primary leaves delayed; dainty foliage; yellowish shoot tips.
grn--granulosa  Base of large, multicellular trichomes greatly enlarged; associated chlorophyll deficiency and lack of vigor may be pleiotropic.
gro--grossa  Cotyledons often more than 2; cotyledons and primary leaf segments short and broad, segments fewer, light grey-green; plants 2/3 normal height, upright, weakly branched; fruit and seed set reduced.
grs--grisea grs1, gr1Smaller, light yellow green plant; dark gray-green growth zones.
gs--green stripe  Irregular longitudinal green stripes in epidermis of unripe fruit; retaininng chlorophyll for longer period during ripening, and eventually assuming paler color in fully ripe fruit; changes limited to epidermis. Striping may be observed on stem under conditions of high humidity and low light; heterozygotes exhibit some faint striping on fruit, but scored as a recessive.
gt--gametophytic factor  Reduced effectivity of female gametes.
Gx--Gibberellin-exserted  Gibberellin-stimulated exsertion of style.
h--hairs absent HLong trichomes absent except on hypocotyl and at growing point; heterozygote is intermediate.
ha--hastata ha1Slender, long, pointed cotyledons; short leaves, with 2 pairs of slender, pointed pinnae; small, flat, dense bush.
he--heteroidea  Small plant; internodes shortened; leaflets somewhat narrowed; older leaves glossy, brittle, and light green.
he-2--heteroidea-2 he2Lax, low bush; slender stems, short internodes; small, keeled pinnae.
Hero--Heterodera rostochiensis resistance  Resistance to potato cyst nematode (Globodera rostochiensis).
hg--heterogemma hg1Small, lax plant; first inflorescence abnormal, numerous fasciated buds, elongated, unbranched peduncle.
hi--hilara  Small plant, proportionately reduced; irregularly crenate and rugose and light-dark green patterned pinnae.
hl--hairless  All types of hairs suppressed except for a few of the glandular type, which lack the chloroplasts normally present; presence of the large multicellular trichome bases imparts a granular or rough appearance to the stems; in contrast to h/h, hypocotyl is glabrous; stamens lacking hairs, tend to be dialytic; plant weak and extremely brittle.
hl2hairless cal, cal1Smaller, weakly pubescent plant; dark-green leaves. Curved upwards to involuted.
hlprov3hairlessprovisional 3hlAll herbage hairless; stems brittle
hlprov4hairlessprovisional 4hlAll herbage hairless; stems brittle
hlprov5hairlessprovisional 5hlAll herbage hairless; stems brittle
hlprov6hairlessprovisional 6hlAll herbage hairless; stems brittle
hl-2--hairless-2 hl^prov6All herbage hairless; stems brittle
hp-1--high pigment-1 hp, hp1, hp2, bs, drChlorophyll, carotenoids, ascorbic acid content of fruit intensified.
hp-1whigh pigment-1WB3 Similar to hp-1, but more extreme phenotype.
hp-2--high pigment-2 hpSimilar to hp-1; enhances all pigments of green and mature red fruits; in presence of u^+ allele, immature fruit are completely dark green.
hp-2dghigh pigment-2dark greendgImmature fruit color normal; darker green color appears as fruit develops, then persists until onset of ripening, high chlorophyll compared to wild type or hp-1.
hp-2jhigh pigment-2JoneshpSimilar to hp-2.
hp-31high pigment-3  Increased accumulation of carotenoids.
Hr--Hirsute  Long hairs on upper leaf surfaces.
Hrt--Hirtum  Increased density of large trichomes; incompletely dominant.
hs--hairless style  Style lack hairs.
ht--hastate  Primary and mature leaves narrow, elongate, and entire, fewer lobed; abortive and deformed flowers.
hy--homogeneous yellow  All vegetative parts yellow in all stages.
I--Immunity to Fusarium wilt race 0  Resistances to Fusarium oxysporum f. lycopersici.
I-2--Immunity to Fusarium wilt race 2  Resistance to Fusarium oxysporum f. lycopersici race 2.
I-3--Immunity to Fusarium race 3  Resistance to Fusarium oxysporum f. lycopersici race 3.
I-4--Immunity to Fusarium wilt-4  Resistance to Fusarium oxysporum race 2.
I-5--Immunity to Fusarium wilt-5  Resistance to Fusarium oxysporum race 2.
I-6--Immunity to Fusarium wilt-6  Resistance to Fusarium oxysporum race 2.
I-7--Immunity to Fusarium wilt-7  Resistance to Fusarium oxysporum race 3.
ic--inclinata  Small bush; short internodes, leaves; older leaves epinastic, yellowish.
ic-2--inclinata-2 ic2Many slender side shoots; short leaves; narrow, smaller pinnae.
ica--icana  Plants reach 3/4 typical size; habit denser with grey-green aspect, revealing some whitish-green of leaf under-surfaces; segments separate with light areas in younger leaves, partly fused, dirty yellow-grey in later leaves.
icn--incana  Small plant, slow growing; whitish seedling and mature leaves latter with prominent purpling.
id--indehiscens  Large pinnae; connate sepals; fruit irregularly cracked. (phenotype similar to cleistogamous).
ida--inordinata  Seedling typical pm; habit erect, irregular bush with less branching; some leaves with 2 pairs of lateral segments, others unsegmented, in later growth becoming normal; term segments almost cordiform; foliolules strongly blistered; foliage color brilliant dark green.
Idh-11Isocitrate dehydrogenase-1   
ig--ignava  Dwarf upright, little branched; leaves relatively large, light green, bleaching to white; growth retarded (1/4 size)
ig-2--ignava-2 ig2Small plant; smaller leaves and pinnae of lighter color; short internodes.
ig-3--ignava-3 ig3Cotyledons small, narrow; lax, flat bush; sturdy shoots, yellowish tips in field; large pinnae.
im--impatiens im1Small, lax plant; reduced side shoots; darker, blue-green foliage wilts in strong sun and wind.
im-2--impatiens-2 im2Small, flat bush; leaves short, small rugose, necrotic at tips; drought sensition.
imb--imbecilla  Plant weak with few branches; younger leaves yellow-green then when older turning light green.
imb-2--imbecilla-2 imb2Light green cotyledons, dark veins; smaller, lax plant; short, yellowish to light green leaves.
impdiaimpedita  Smaller, weak plant; leaves chlorotic at margins; plant smaller than imp:eg.
impegimpedita  Smaller, weak plant; leaves chlorotic at margins, like imp;dia but plant larger.
in-2--indiga-2 in2Narrow, gray-green cotyledons; smaller plant; short internodes.
in-3--indiga-3 in3Small plant; smaller pinnae; reduced branching; inflorescences close together and numerous.
ina--inflexa ina1Lax growth, becoming partially decumbent.
inc--incurva  Leaf segments weakly curved, undulate margins; stems, leaf veins and inflorescence crooked.
inc-2--incurva-2 inc2Smaller plant; short internodes and leaves; long, narrow, keeled and brownish-green pinnae.
inf--informa  Tiny plant with very short internodes; leaves small, plicate, and strongly down-curved.
ini--inquieta ini1Small, proportionately reduced, variable plant; young pinnae involuted, lighter colored; large trichomes absent.
ino--involuta ino1Smaller, variable, cylindrical bush; short internodes; involuted pinnae.
inr--integer inr1Primary leaves almost undivided; long petioles; broad, rugose pinnae; few secondary pinnae.
ins--inconstans ins1Small, weakly branched plant; phenotype, including leaf size, variable from year to year.
inta--integrifolia inta1Lax, high bush; short leaves; rounded, crowded pinnae.
inv--invalida  Small, dainty plants; irregular chlorotic flecks, later leaves deformed.
inx-2--introflexa-2 inx2, intro2Plants diminutive in all respects; keeled to rolled, wilty pinnae.
Ip--Intense pigment  Dark pigmentation of the unripe fruit.
irr--irregularis  Leaves shortened, irregularly veined, dark green, tinged yellow. Leaves erect on stem and stems swollen at leaf nodes.
ita--inquinata ita1Young leaves in midseason develop small necrotic flecks, later coalescing into large ones; variable expressivity.
Itm--Intumescences  Development of tumorous outgrowths on leaves and stems; heterozygote intermediate.
j--jointless lfThe normal pedicel joint, with swelling and abscission layer is absent; fruits separate at the juncture with the calyx; leafy or indeterminate growth of the inflorescence, suppressed in combination with sp (self-pruning).
j-2--jointless-2 j2Jointless elongate pedicel; teratological calyx; proliferated inflorescence; elongate fruit.
j-2injointless-2 j2^inJointless pedicel, with incomplete gene action. Pedicel joints with normal form, but do not separate when fruits ripen. Referred to as the 'arthritic' gene.
Jau--Jaundiced  Yellow-green seedlings and young plants; homozygous lethal
jug--jugata  Fasciated stems and fruits; short internodes; inflorescence almost sterile.
jug2jugata jug1^2Fasciated stems, flowers, fruits, more extreme than jug.
l--lutescent gCotyledons and first true leaves turn yellow when the seedlingsa re 4-6 weeks old; as plants grow, the older leaves progressively yellow and abscise prematurely, the growing tips and immature leaves remaining green; stigmas yellow and fruit pigmentation reduced.
l2lutescent rubYellowish unripe fruits; premature yellowing of the leaves.
lprov3lutescentprovisional 3lYellowish unripe fruits; premature yellowing of the leaves.
lprov4lutescentprovisional 4lYellowish unripe fruits; premature yellowing of the leaves.
l-2--lutescent-2 l-3, l2Premature yellowing of older leaves; yellowish unripe fruit.
La--Lanceolate  Leaves simple, limited to one small, elongated leaflet; margins entire; stems slender and excessively branched; cotyledons frequently fused; fruit small; homozygous inviable (shoot growth normally produces only a hypocotyl with little if any leaf tissue).
Lac--Lacciniate  Leaf margins deeply incised, segments narrow.
lae--laesa  Small bush; axillary sprouts later necrotic; primary leaves dainty.
lan--languida  Plant size and habit typical pm; until fruiting, variably light green virescent, later typical pm.
lap--lamprochlora lap1Small, lax plant; short, shiny leaves; end pinnae curved upwards.
lat--lata  Habit broad at first; growth much depressed in field.
lax--laxus lax1Plant lax, weakly branched; slender shoots; smaller, slightly rounded pinnae.
Lc--Locule number  Locule number is reduced; associated with o.
le--lembiformis le1Prostrate, smaller plant, proportionately reduced; keeled or involuted yellowish pinnae, ventrally purplish.
lep--leprosa lep1Smaller, proportionately reduced plant; rapidly spreading necrosis on all older pinnae.
lg--light-green lmeLight green foliage.
lg-2--light green-2 lg2, li, pg392Cotyledons light yellow; leaves pale green; classification sometimes difficult.
lg-3--light green-3 lg3, lt, pg483Light green cotyledons, leaves; former fade to yellow, latter to pale-green.
lg-4--light green-4 lg4Yellow-green cotyledons, light green leaves.
lg-5--light green-5 lg5, lm, fy, ytFoliage uniformly light green
li--limbrata  Seedling development variably reduced; first leaves slightly deformed with terminal segments broader at tip, in early stages pinnae tend to be finer, more strongly plicate, margins curled upward, and yellow-green. These characters disappear in mature plant.
liv--livida  Plant erect, near normal size; cotyledons narrow, light grey-green, white-spotted; leaf segments long, narrow, slightly notched, blunt, dull grey-green; shoot tips bluish with anthocyanin.
Ln--Lanata  Resembles Wo:m but hairs concentrated on stem; all 3 genotypes viable and distinguishable.
LnGLanata  resembles Wo:m but is more hairy; all 3 genotypes viable and distinguishable.
lo--locule number reduced  Reduced locule number.
loa--low acid  Low titratable fruit acidity.
loc--locule number increased  Locule number increased.
lop--longipes lop1Small, somewhat spreading plant; broad, weak gray-green pinnae; long petioles; variable expression.
Lpg--Lapageria  Leaves small, dark green, glossy; flowers campanulate; homozygote sterile.
ls--lateral suppresser  Few or no axillary branches; corolla suppressed; partially male sterile
ls2lateral suppresser  Few or no axillary branches; corolla suppressed; partially male sterile.
lt--laeta lt1Smaller, weakly branched plant; light gray-green foliage; growing points, later entire plant, yellow-green.
lt-2--laeta-2 lt2Dainty, light green foliage; shoot tips with yellowish pinnae and in late season has bluish veins, petioles.
ltf--latifolia  Cotyledons and pinnae extremely broad proportioned.
lu--luteola  Light yellow corolla. .
luc--lucida  Leaves bright green, yellow-green at growing point.
luc-2--lucida-2 luc2Smaller, lax bush; short stems; small, light green leaves.
luc-3--lucida-3 luc3Plant and leaves small, light green to light olive.
lur--lurida lur1Yellow-green cotelydons and primary leaves; dull light green later foliage.
lut--lutea  Leaves yellow-green, darker veins; virescent.
lut-2--lutea-2 lut2Primary leaves flecked yellow-green; pinnae dull light green, yellowing early.
lut-3--lutea-3 lut3Cotyledons yellowish light green, small plant and leaves of dull light green.
lux--luxurians lux1Young plant larger, fewer side shoots; erect, long internodes.
Lv--Leveillula taurica resistance  Resistance to Leveillula taurica powdery mildew.
Lx--Lax  Elongate, acute-segmented, pendent leaves.
lyr--lyrate  First leaves entire and undivided; later fan shaped and deeply lobed; female-sterile. Notes: calyx greatly enlarged; heterozygotes have the leaf phenotype but not sterility.
lz--lazy  Normal seedlings, 3 weeks later become strongly geotropic; weak branching.
lz-2--lazy-2  Plant habit strongly prostrate due to reversed geotropic response of shoots (roots normal).
m--mottled  Cotyledons and all leaves speckled with light yellow-green spots, rather evenly distributed; leaflets narrow, irregular and distorted.
m-2--mottled-2 m2, mo, mdMany small chlorotic spots on leaves; plant stunted; temperature-dependent.
M-3--Mottled-3 M3, M345Variable sectoring for chlorophyll deficiency; homozygous lethal.
ma--macrocarpa  Leaves clearly longer; fruit large, smooth.
maa--macra  Branching inhibited, erect habit; leaf segments and flowers reduced size.
mac--maculata mac1Small plant; necrosis on upper or lower leaf surface spreading from margin.
mad--marcida mad1Small plant; dark-green, slightly wilty foliage.
Mae-11Malic enzyme-1  Allele of S. lycopersicoides, slightly retarded from '+'.
mal--maculosa mal1Young plant leaves have irregular, yellowish light green flecks, disappear later.
mar--marcescens  Small plant; leaves dark-green, lightly wilting.
marm--marmorata  Marbling of several tones of white and light green on leaves, strongest expression in greenhouse.
marm2marmorata marm1^2Marbling of several tones of white and light green on leaves, strongest expression in greenhouse.
marm-2--marmorata-2  Cotyledons variably white, yellow, yellow-green or green with variegated true leaves.
mat--matura mat1Lax, flat bush, retarded in growth; short leaves; half-sized pinnae.
mc--macrocalyx  Calyx variable in size, but always larger than normal, exceeding corolla in length and often bladdery in appearance; corolla subnormal in development and rarely spread at anthesis; pedicel joint arthritic; inflorescence indeterminant.
mcn--maculonecrotic  Seedling 2/3 normal size with virescence, yellow spots becoming necrotic, mature leaves often deformed, especially mid-region, also virescent at maturity.
mcr--multicolor  Seedling typical spl; plant 3/4 size; habit spreading; foliage color yellow green early to dark midveins with necrotic spots later, finally with variable whitish-yellow-green aspect.
mcs--macrosepala  Size and habit normal; inflorescence elongate, indeterminate; corolla enlarged; closed, but opening when touched; calyx with similar features; older leaves uncurled to reveal undersides. Note: calyx segments fused.
Mdh-11Malate dehydrogenase-1   
Mdh-21Malate dehydrogenase-2   
Mdh-41Malate dehydrogenase-4   
Me--Mouse ears  Leaves 3-4 pinnately compound with clavate segments; shortened internodes; homozygous viable; incompletely dominant.
med--mediocris med1Smaller plant, proportionately reduced.
mel--melongenoida mel1Sometimes lax habit; oval fruit.
men--mendosa  Seedlings variable size; plant reaches half normal size; leaves always deformed, segments reduced, curled, very blistered; flowers deformed, style exserted from bud stage.
mes--methyl salicylate concentration  High levels of methyl salicylate in fruit of certain lives.
mf--multiformis  Growth always retarded; thin branches; internodes and leaves shortened, segments small, oval, almost entire; very few foliolules, foliage starts grey-green, later dull green.
mgn--marginal necrotic  Seedling 1/4 normal size; cotyledons and leaves yellowish, becoming necrotic first at tips and margins; mature plant subnormal in size and leaves with brown necrotic blotches.
Mi--Meloidogyne incognita resistance  High level resistance to root-knot nematode species, including M. incognita, M. arenaria, and M. javanica. Resistance breaks down at soil temperatures above 28 deg. C. Also confers resistance to potato aphid and whitefly.
Mi-3--Meloidogyne incognita resistance-3  Resistance to Mi-breaking strains of Meloidogyne incognita, including at high temperatures.
Mi-4--Meloidogyne incognita resistance-4