List of Gene Names and Symbols
Gene Allele Gene Name Allele Name Synonyms of Allele Phenotype
(G)




(K) -- (K)

Inhibits modifiers; permitting expression of tangerine gene; interacts with yellow fruit flesh gene.
a -- anthocyaninless
a1 Stems and leaves always lack anthocyanin.
a prov2 anthocyaninless
a Stems and leaves always lack anthocyanin.
a prov3 anthocyaninless
a Stems and leaves always lack anthocyanin.
aa -- anthocyanin absent

Complete absence of anthocyanin in all parts
Abg -- Aubergine

Fruit epidermis purple, particularly on shoulder and where exposed to direct light; also enhanced by wounding.
abi -- aborted inflorescence

Inflorescences reduced to an abortive 2-3 mm structure.
ac -- apocarpous

Highly deformed multicarpellate and apocarpous fruits
Aco-1 2 Aconitase-1


Aco-1 3 Aconitase-1


Aco-1 1 Aconitase-1


Aco-2 2 Aconitase-2


Aco-2 1 Aconitase-2


acr -- acroxantha
acr1 Smaller plant; short internodes; rugose, dull, light green leaves, turn yellow starting at tip.
acu-2 -- acumbens-2
acu2 Short, rounded cotyledons; short, compacted leaves.
ad -- Alternaria alternata resistance

Resistance to Alternaria collar rot.
Adh-1 1 Alcohol dehydrogenase-1


Adh-1 2 Alcohol dehydrogenase-1


Adh-1 n Alcohol dehydrogenase-1


Adh-2 1 Alcohol dehydrogenase-2


adp -- adpressa

Plant small and decumbent with short internodes and leaves.
adp-2 -- adpressa-2
adp2 Prostrate plant habit; short internodes and leaves; rugose, bright green pinnae.
adu -- adusta
adu1 Small plants, proportionately reduced; progressive necrosis from older to younger leaves.
adu-2 -- adusta-2
adu2 Stems and leaves shortened, develop progressive necrosis; small, rounded, rugose pinnae
adu-3 -- adusta-3
adu3 Slightly smaller leaves, shiny above, develop progressive necrosis.
ae -- entirely anthocyaninless
a332 Completely free of anthocyanin.
ae 2 entirely anthocyaninless

Completely free of anthocyanin.
ae afr entirely anthocyaninless fragilis afr, ap Anthocyanin absent; small, brittle, fragile plants; leaves wilt in daytime.
ae prov3 entirely anthocyaninless
ae Lacks anthocyanin in the seedling stem.
aeg -- aegrota

Brown necrotic layer spreads over leaves, which die prematurely.
aeg-2 -- aegrota-2
aeg2 Dainty leaves turn yellow and die early.
aer -- aerial roots

Adventitious roots on stem from soil level to considerable height above.
aer-2 -- aerial roots-2

Abundance of root initials along the stems.
af -- anthocyanin free
a325 Completely free of anthocyanin; lacks glandular hairs.
Af. -- Anthocyanin fruit

Anthocyanin in green and ripe fruit; environmentally sensitive, absent when shaded
afe -- afertilis
afe1 Small plant; short internodes, leaves; dainty, yellowish, mostly keeled pinnae; poor fruit set; heterozygote intermediate.
afl -- albifolium
af Seedling very slow; cotyledons white; leaves becoming normal except for gray flecks; later growth nearly normal.
afl-2 -- albifolium-2

Cotyledons whitish light green; pinnae with irregular yellow-white areas; green-white stripes on the stems and petioles; foliage of mature plant grey-green with whitish blotches; plants erect, 1/3 - 2/5 normal size. Slightly dominant, the heterozygote attaining 2/3 -3/4 normal size.
ag (2k) anthocyanin gainer Knowles 2nd ag^4 Anthocyanin absent except on cotyledons and lower sides of leaves.
ag -- anthocyanin gainer

Anthocyanin absent except on cotyledons and lower sides of leaves.
ag (1k) anthocyanin gainer Knowles ag^3 Anthocyanin absent except on cotyledons and lower sides of leaves. (Indistinguishable from ag).
ag (1s) anthocyanin gainer Stubbe's ag^2 Anthocyanin absent except on cotyledons and lower sides of leaves.
ag 2 anthocyanin gainer

Completely free of anthocyanin.
ag k anthocyanin gainer Kosala's
As for ag, purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
ag-2 -- anthocyanin gainer-2

As for ag purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
ags-2 -- aegrescens-2

Plant 1/3 normal size; reduced branching; leaves shortened; pinnae small, olive green, becoming necrotic at tips.
ah prov2 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah -- Hoffman's anthocyaninless
ao, a337 Completely free of anthocyanin
ah prov7 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov6 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov5 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov4 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov3 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ai -- incomplete anthocyanin
a342 Early seedling stem has trace of anthocyanin, soon lost.
ai 2 incomplete anthocyanin
am, a340 Early seedling stem has trace of anthocyanin, soon lost. (Typical green stem mutant.)
al pu-2 anthocyanin loser pulvinata-2 pu-2 Semispherical bush, anthocyaninless; short internodes.
al -- anthocyanin loser
a2 Purple stems become green in 10 to 21 days.
al-2 -- anthocyanin loser-2

No anthocyanin in cotyledons and primary leaves; little anthocyanin in hypocotyl; internodes and leaves shorted; pinnae and interpinnae modified.
ala -- albina
ala1, alb1 White or cream-colored cotyledons.
ala-2 -- albina-2
alb2, ala2 White or cream-colored cotyledons.
ala-3 -- albina-3
ala3, alb3 White or cream-colored cotyledons.
ala-4 -- albina-4
ala4, alb4 White or cream-colored cotyledons; lethal.
alb -- albescent

White variegation promoted by low temperatures.
alb prov2 albescent
alb Same as alb.
alc -- alcobaca

Greatly delayed fruit ripening processes.
alu -- alutacea
alu1 Small plant, yellowish foliage.
alv -- alvoviridis
alb, alb1 Whitish green cotyledons, pinnae; strong anthocyanin in stems, veins.
Am -- Alfalfa mosaic

Resistance to alfalfa mosaic virus
ama -- albomarginata

Shorter internodes; leaves small and spotted with yellow-green or yellow; flowers small and petals narrow with white edges.
an -- anantha
an^1, an^2, ca Flowers greatly modified; inflorescences compound; closely resembles ca.
an-2 -- anantha-2

Inflorescence highly branched, terminates in very compact ovary-like masses of tissue; aborted flowers.
an-3 -- anantha-3

Leaves small, light green; inflorescence indefinitely ramifying, no normal flowers, resembling cauliflower; completely sterile.
ana -- angustata
ana1 Shortened leaves, stems; pinnae small, narrow, pointed; smaller flowers, narrow petals.
ang-2 -- angustifolia-2
ang2 Smaller, compact, slow-growing plant; narrow, light-green pinnae.
anr -- antirrhinum

Fruit elongated with hollow tip from which epiglotis extends resembling a snapdragon flower.
ant -- aurantia
ant1 Short thick stems; many short side shoots; narrow, rugose, light green pinnae; light orange fruit; colorless pericarp.
anu -- angusta
anu1 Dark gray-green pinnae narrowed to 1/3 normal width; short leaves.
ap -- apetalous

Corolla reduced or absent; pollen scarce and nonfunctional
apl -- applanata

Broad, spreading habit; long, laxly pinnate leaves; heterozygote intermediate.
apn -- albo-punctata

Fine white speckling uniformly on cotelydons; mature leaves small, wilty; tiny flowers.
Aps-1 1 Acid phosphatase-1


Aps-1 2 Acid phosphatase-1


Aps-1 n Acid phosphatase-1


Aps-2 1 Acid phosphatase-2


Aps-2 2 Acid phosphatase-2


Aps-2 3 Acid phosphatase-2


Aps-2 n Acid phosphatase-2


ar -- arrecta

Small, compact plants; leaves yellow-green, virescent.
are -- anthocyanin reduced

Almost no anthocyanin until fruits set, then moderate amount in young foliage.
ari -- aridifolia

Plants 1/3 normal size; cotyledons small, pointed, dull dark grey-green; die prematurely.
as -- asynaptic
as1 Asynaptic meiosis; high pollen and ovule sterility.
as-2 -- asynaptic-2
as2 Asynaptic meiosis; high pollen and ovule sterility.
as-3 -- asynaptic-3
as3 Asynaptic meiosis; high pollen and ovule sterility.
as-4 -- asynaptic-4
as4 Asynaptic meiosis; high pollen and ovule sterility.
as-5 -- asynaptic
as5 asynaptic meiosis; high pollen and ovule sterility.
as-6 -- asynaptic-6
as6 Delayed and asynaptic meiosis; chromatin degeneration; possibly two loci involve
Asc -- Alternaria stem canker resistance

Resistance to Alternia stem canker
at -- apricot

Yellow-pink color of fruit flesh
atn -- attenuata
at Narrow, pointed leaves, yellow-green, purple below; rigid cylindrical habit; thin shoots.
atv -- atroviolacium

Excess anthocyanin on leaves, stems, and fruits.
au 6 aurea
yg^6, yg-6, au^yg-6, yo Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
au (1k) aurea Knowles

au -- aurea

Bright yellow cotyledons and leaves; seedling etiolates; plant vigorous.
au tl aurea Torrey Lyons'
Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
au 3 aurea
au-2, au2 Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au 2 aurea
au^3 Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au (2k) aurea Knowles second

au (1s) aurea Stubbe's au^2, au, brac Bright yellow cotyledons and leaves; some times whitish; corolla pale yellow; moderately vigorous in condine red; seedling etiolates; plant vigorous.
au w aurea
w616 Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
auc -- aucta

Few branches; leaves broad, rounded, later becoming blistered, dark green; number of flowers and fruits reduced.
aur -- aurantiaca
aur1 Small, pointed, yellowish light-green pinnae; orange fruit.
aus -- austera

Plants always 1/3 to 2/3 normal size; pinnae finely divided, in some cases bent upwards; foliage darker and duller than normal.
aut -- aureata

Vireescent, flush of gold near growing point in early development, not detectable in older plants.
auv -- aureate virescent

Seedling 'gold dust' virescence; leaf segments pointed, sometimes fimbriate. Mature plant nearly normal.
avi -- albovirens
avi1 Smaller plant; foliage variously variegated yellow, white, grey-green; reduced fruit set.
aw prov4 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw prov5 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw prov3 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw -- without anthocyanin
aba, ab, a179 Free of anthocyanin, same phenotype as a.
aw (1k) without anthocyanin
aw^2 Completely free of anthocynanin. Radiation induced in Money Maker.
aw-2 -- without anthocyanin-2

No anthocyanin at any stages; reduced growth rate, dense bushy habit.
B -- Beta-carotene

High B-carotene, low lycopene in ripe fruit.
B og Beta-carotene old gold og Corolla tawny orange; increased fruit lycopene.
B m Beta-carotene minutum
High B-carotene, low lycopene in ripe fruit. .
B c Beta-carotene crimson og^c,Crn,Cr,crn-2,cr-2 Increased fruit lycopene content; phenotype similar to B^og.
bc -- bicolor
bi Young leaves mottled brownish olive normal, later glazed dark green; short internodes; heterozygote intermediate for some characters.
Bco -- Brilliant corolla

Corolla pigment intensifier.
be -- blue-green

Small plant with blue-green leaves.
Ber
Blossom-end rot resistance


bg -- bursting resistance

Resistance of fruit to bursting.
bi -- bifurcate inflorescence

With f and j causes extreme fasciation of the "ring type".
bip -- bipinnata

Leaves highly divided, with smaller, acute segments.
bip prov2 bipinnata
bip Leaves highly divided, with smaller, acute segments.
bip-2 -- bipinnata-2
bip2 Slender, pointed, light gray-green cotyledons; doubly pinnate, rugose leaves.
bk -- beaked

Fruits with sharp points at stylar end.
bk 2 beaked

Fruits with sharp points at stylar end.
bk 3 beaked

Fruits with sharp points at stylar end.
Bk-2 -- Beaked-2

Sharp point at stylar end of fruit
bl -- blind

Stem terminating in first inflorescence; midribs may develop adventitious shoots.
bl 2 blind
to^2 Determinant growth; few flowered inflorescences; fasciated flowers; enlarged, leafy calxy; less extreme than bl.
bl to blind torosa to Determinate habit; firm, strongly uprolled pinnae; fasciated flowers and fruits.
bls prov2 baby lea syndrome
bls Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bls -- baby lea syndrome
alm Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bls prov3 baby lea syndrome
bls Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bn -- blunt

Stamens absent, rarely present, with functional pollen, in greenhouse. Usually petaloid stamens adnate to pistil; leaf segments obtuse
bn sts blunt Stamen suppressor sts, sl-6 Stamens absent, rarely present, with functional pollen, in greenhouse; fewer blunt leaflets; with sp:+ 6-10 leafs between inflorescences.
Bnag-1 1 Beta-N-acetyl-D-glucosaminidase-1


br -- brachytic

Internodes shortened
brt -- bushy root

Radical branches early; radical and root profusely branched.
brt-2 -- bushy root-2

severely stunted growth (1/20) dense bushy growth of twisted roots.
bs -- brown seed

Brown seed color behaving as endosperm trait.
bs-2 -- brown seed-2

Brown seed color governed by genotype of endosperm.
bs-3 -- brown seed-3

Brown seed color determined by genotype of endosperm.
bs-4 -- brown seed-4

Brown seed color controlled by genotype of endosperm.
Bt -- bursting digenic inheritance
bg Susceptibility to fruit bursting.
btl -- brittle stem

Growth ceases with first flowering; stems brittle, with progressive defoliation; temperature-sensitive and expressed only under field conditions; B inefficient.
bu cin bushy compact inflorescence cin Internodes and inflorescences shortened; long petioles.
bu ab bushy abbreviata fru^ab Internodes and inflorescences shortened; long petioles.
bu cin-2 bushy compact inflorescence 2 cin-2 Internodes and inflorescences shortened; long petioles. Duplicate genes determining character similar to bu.
bu hem bushy hemiglobosa fru^hem Internodes and inflorescences shortened; long petioles. Phenotype like bu except leaves longer, more lax; heterozygote intermediate in some traits.
bu -- bushy
fru Internodes and inflorescences shortened; long petioles.
bul -- bullata

Leaf surface blistered and chlorotic with a network of darker veins; short internodes.
buo -- bullosa
buo1 Plants and seedlings larger in all parts; broad and rugose pinnae.
c prov3 potato leaf
c Number of leaf segments reduced
c prov4 potato leaf
c Number of leaf segments reduced
c prov6 potato leaf
c Number of leaf segments reduced
c prov2 potato leaf
c Number of leaf segments reduced
c prov5 potato leaf
c Number of leaf segments reduced
c -- potato leaf

Number of leaf segments reduced
c int potato leaf integerrima int Number of leaf segments reduced
car -- carinata

Shortened, keeled pinnae, dark gray-green color; deformed flowers; elongated fruit.
car-2 -- carinata-2
car2 Smaller plant; slender stems; smaller pinnae, nearly entire margin.
cb -- cabbage

Compact habit; leaves large and dark green; fewer locules.
cb-2 -- cabbage leaf-2

Larger plant parts; leaves epinastic and blistered.
cc
constricting corolla


ccf -- cactiflora

Flower greatly fasciated; and roecium gynoecium and green tissues fused and distorted, enlarging into fruitlike structure; complete make and female sterility.
cd -- condensata

Seedling small; internodes shortened; leaf light grey-green with narrow, keeled segments; fruit round and relatively large.
ce -- cernua
ce1 Cotyledons lightly dotted in whitish yellow-green; lax, inclined plant.
Cf-? -- Cladosporium fulvum resistance

Resistance to specific races of Cladosporium fulvum
Cf-1 -- Cladosporium fulvum resistance-1
Cf, Cf1, Cfsc Resistance to race 1 and 3 of Cladosporium fulvum.
Cf-1 3 Cladosporium fulvum resistance-1
Cf-5, Cf5 Resistance to specific races of Cladosporium fulvum.
Cf-10
Cladosporium fulvum resistance-10


Cf-12
Cladosporium fulvum resistance-12


Cf-19
Cladosporium fulvum resistance-19


Cf-2 -- Cladosporium fulvum resistance-2
Cf2, Cfp1 Resistance to races of Cladosporium fulvum.
Cf-3 -- Cladosporium fulvum resistance-3
Cf3, Cfp2 Resistance to specific races of Cladosporium fulvum
Cf-4 -- Cladosporium fulvum resistance-4
Cf-8, Cf4, Cf-1^2 Resistance to specific races of Cladosporium fulvum.
Cf-5
Cladosporium fulvum resistance-5


Cf-6 -- Cladosporium fulvum resistance-6

Resistance to specific races of Cladosporium fulvum.
Cf-7 -- Cladosporium fulvum resistance-7

Resistance to specific races of Cladosporium fulvum.
Cf-9 -- Cladosporium fulvum resistance-9

Resistance to specific races of Cladosporium fulvum.
cfa -- conferta
cfa1 Very small, compact bush; strongly shortened internodes, young leaves.
cg -- congesta
cg1 Small, dense bush; late growth retardation; short internodes; pointed primary leaves.
ch -- chartreuse

Greenish-yellow subnormal petals; reduced fertility.
chln -- chloronerva

Upper leaves smaller and chlorotic interveinally; strong graft response.
chrs -- chrysanthemum sterile

Flowers fascicled 6cm across; many small pale yellow petals; stamens dialytic and without pollen; fruits deformed, orange-red fruits with brown sectors; poor fertility.
ci -- cincta
ci1 Small, irregular bush; many side shoots of equal length from lower leaf axils.
cin-2
compact inflorescence-2


cir -- circumsaepta
cir1 Rounded, light gray-green pinnae, downrolled margin.
cit -- citriformis

In pm stock; Plant reaches 1/3 - 2/5 typical size; half erect, weakly branched; foliage color grey-green; terminal segment of primary leaves long and narrow; older leaves serrated; lemon shaped fruit with conspicuous stylar scar.
cjf -- confunctiflora

Flowers joined in pairs or triplets, otherwise malformed.
ck -- corky fruit

Pericarp cracked, exposed tissues becoming brown and leathery.
Cl -- Cleistogamous
Cl1 Flowers fail to open.
cl-2 -- cleistogamous-2
cl2 Flowers open only slightly; reduced fertility.
cla -- clara

Yellowish-green leaf color, purple veins and petioles; incompletely dominant.
clau ics clausa incisifolia ics Pinnae margins deeply incised, dentations acute.
clau prov2 clausa
clau Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau ff clausa

Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau -- clausa
ff, vc Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau vc clausa

Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
cle -- collecta

Reduced growth rate; internodes shortened; small compact bush habit; leaf segments large, rounded, foliolules fewer.
cls -- clarescens

Foliage light yellow green early, becoming whitish green and blistered later, brilliant yellow green virescent; plants 1/4 typical size; growth very weak in field, setting fruits only in greenhouse.
clt -- coalita

Size and habit typically spl; primary leaves with entire end segments and only tiny laterals; mature leaves typically 5-segmented through fusion with end segment; segments always entire; resembles but is not allelic to sf.
clt-2 -- coalita-2

Terminal pinnae large, second order pinnae few and masked by 1st order pinnae, little serration.
cm -- curly mottled

Strong virus-like mottling and distortion of leaves and abortion of flowers; highly sensitive to environment.
cma -- commutata

dwarf with bright green foliage
Cmm -- Clavibacter michiganensis resistance

Resistance to Clavibacter michiganensis.
Cmr -- Cucumber mosaic resistance