List of Gene Names and Symbols
Gene Allele Gene Name Allele Name Synonyms of Allele Phenotype

(K) -- (K)

Inhibits modifiers; permitting expression of tangerine gene; interacts with yellow fruit flesh gene.
a -- anthocyaninless
a1 Stems and leaves always lack anthocyanin.
a prov2 anthocyaninless
a Stems and leaves always lack anthocyanin.
a prov3 anthocyaninless
a Stems and leaves always lack anthocyanin.
aa -- anthocyanin absent

Complete absence of anthocyanin in all parts
Abg -- Aubergine

Fruit epidermis purple, particularly on shoulder and where exposed to direct light; also enhanced by wounding.
abi -- aborted inflorescence

Inflorescences reduced to an abortive 2-3 mm structure.
ac -- apocarpous

Highly deformed multicarpellate and apocarpous fruits
Aco-1 2 Aconitase-1

Aco-1 3 Aconitase-1

Aco-1 1 Aconitase-1

Aco-2 2 Aconitase-2

Aco-2 1 Aconitase-2

acr -- acroxantha
acr1 Smaller plant; short internodes; rugose, dull, light green leaves, turn yellow starting at tip.
acu-2 -- acumbens-2
acu2 Short, rounded cotyledons; short, compacted leaves.
ad -- Alternaria alternata resistance

Resistance to Alternaria collar rot.
Adh-1 1 Alcohol dehydrogenase-1

Adh-1 2 Alcohol dehydrogenase-1

Adh-1 n Alcohol dehydrogenase-1

Adh-2 1 Alcohol dehydrogenase-2

adp -- adpressa

Plant small and decumbent with short internodes and leaves.
adp-2 -- adpressa-2
adp2 Prostrate plant habit; short internodes and leaves; rugose, bright green pinnae.
adu -- adusta
adu1 Small plants, proportionately reduced; progressive necrosis from older to younger leaves.
adu-2 -- adusta-2
adu2 Stems and leaves shortened, develop progressive necrosis; small, rounded, rugose pinnae
adu-3 -- adusta-3
adu3 Slightly smaller leaves, shiny above, develop progressive necrosis.
ae -- entirely anthocyaninless
a332 Completely free of anthocyanin.
ae 2 entirely anthocyaninless

Completely free of anthocyanin.
ae afr entirely anthocyaninless fragilis afr, ap Anthocyanin absent; small, brittle, fragile plants; leaves wilt in daytime.
ae prov3 entirely anthocyaninless
ae Lacks anthocyanin in the seedling stem.
aeg -- aegrota

Brown necrotic layer spreads over leaves, which die prematurely.
aeg-2 -- aegrota-2
aeg2 Dainty leaves turn yellow and die early.
aer -- aerial roots

Adventitious roots on stem from soil level to considerable height above.
aer-2 -- aerial roots-2

Abundance of root initials along the stems.
af -- anthocyanin free
a325 Completely free of anthocyanin; lacks glandular hairs.
Af. -- Anthocyanin fruit

Anthocyanin in green and ripe fruit; environmentally sensitive, absent when shaded
afe -- afertilis
afe1 Small plant; short internodes, leaves; dainty, yellowish, mostly keeled pinnae; poor fruit set; heterozygote intermediate.
afl -- albifolium
af Seedling very slow; cotyledons white; leaves becoming normal except for gray flecks; later growth nearly normal.
afl-2 -- albifolium-2

Cotyledons whitish light green; pinnae with irregular yellow-white areas; green-white stripes on the stems and petioles; foliage of mature plant grey-green with whitish blotches; plants erect, 1/3 - 2/5 normal size. Slightly dominant, the heterozygote attaining 2/3 -3/4 normal size.
ag (2k) anthocyanin gainer Knowles 2nd ag^4 Anthocyanin absent except on cotyledons and lower sides of leaves.
ag -- anthocyanin gainer

Anthocyanin absent except on cotyledons and lower sides of leaves.
ag (1k) anthocyanin gainer Knowles ag^3 Anthocyanin absent except on cotyledons and lower sides of leaves. (Indistinguishable from ag).
ag (1s) anthocyanin gainer Stubbe's ag^2 Anthocyanin absent except on cotyledons and lower sides of leaves.
ag 2 anthocyanin gainer

Completely free of anthocyanin.
ag k anthocyanin gainer Kosala's
As for ag, purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
ag-2 -- anthocyanin gainer-2

As for ag purple pigment appears on cotyledons and lower sides of leaves when growth is slow.
ags-2 -- aegrescens-2

Plant 1/3 normal size; reduced branching; leaves shortened; pinnae small, olive green, becoming necrotic at tips.
ah prov2 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah -- Hoffman's anthocyaninless
ao, a337 Completely free of anthocyanin
ah prov7 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov6 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov5 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov4 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ah prov3 Hoffman's anthocyaninless
ah Completely free of anthocyanin
ai -- incomplete anthocyanin
a342 Early seedling stem has trace of anthocyanin, soon lost.
ai 2 incomplete anthocyanin
am, a340 Early seedling stem has trace of anthocyanin, soon lost. (Typical green stem mutant.)
al pu-2 anthocyanin loser pulvinata-2 pu-2 Semispherical bush, anthocyaninless; short internodes.
al -- anthocyanin loser
a2 Purple stems become green in 10 to 21 days.
al-2 -- anthocyanin loser-2

No anthocyanin in cotyledons and primary leaves; little anthocyanin in hypocotyl; internodes and leaves shorted; pinnae and interpinnae modified.
ala -- albina
ala1, alb1 White or cream-colored cotyledons.
ala-2 -- albina-2
alb2, ala2 White or cream-colored cotyledons.
ala-3 -- albina-3
ala3, alb3 White or cream-colored cotyledons.
ala-4 -- albina-4
ala4, alb4 White or cream-colored cotyledons; lethal.
alb -- albescent

White variegation promoted by low temperatures.
alb prov2 albescent
alb Same as alb.
alc -- alcobaca

Greatly delayed fruit ripening processes.
alu -- alutacea
alu1 Small plant, yellowish foliage.
alv -- alvoviridis
alb, alb1 Whitish green cotyledons, pinnae; strong anthocyanin in stems, veins.
Am -- Alfalfa mosaic

Resistance to alfalfa mosaic virus
ama -- albomarginata

Shorter internodes; leaves small and spotted with yellow-green or yellow; flowers small and petals narrow with white edges.
an -- anantha
an^1, an^2, ca Flowers greatly modified; inflorescences compound; closely resembles ca.
an-2 -- anantha-2

Inflorescence highly branched, terminates in very compact ovary-like masses of tissue; aborted flowers.
an-3 -- anantha-3

Leaves small, light green; inflorescence indefinitely ramifying, no normal flowers, resembling cauliflower; completely sterile.
ana -- angustata
ana1 Shortened leaves, stems; pinnae small, narrow, pointed; smaller flowers, narrow petals.
ang-2 -- angustifolia-2
ang2 Smaller, compact, slow-growing plant; narrow, light-green pinnae.
anr -- antirrhinum

Fruit elongated with hollow tip from which epiglotis extends resembling a snapdragon flower.
ant -- aurantia
ant1 Short thick stems; many short side shoots; narrow, rugose, light green pinnae; light orange fruit; colorless pericarp.
anu -- angusta
anu1 Dark gray-green pinnae narrowed to 1/3 normal width; short leaves.
ap -- apetalous

Corolla reduced or absent; pollen scarce and nonfunctional
apl -- applanata

Broad, spreading habit; long, laxly pinnate leaves; heterozygote intermediate.
apn -- albo-punctata

Fine white speckling uniformly on cotelydons; mature leaves small, wilty; tiny flowers.
Aps-1 1 Acid phosphatase-1

Aps-1 2 Acid phosphatase-1

Aps-1 n Acid phosphatase-1

Aps-2 1 Acid phosphatase-2

Aps-2 2 Acid phosphatase-2

Aps-2 3 Acid phosphatase-2

Aps-2 n Acid phosphatase-2

ar -- arrecta

Small, compact plants; leaves yellow-green, virescent.
are -- anthocyanin reduced

Almost no anthocyanin until fruits set, then moderate amount in young foliage.
ari -- aridifolia

Plants 1/3 normal size; cotyledons small, pointed, dull dark grey-green; die prematurely.
as -- asynaptic
as1 Asynaptic meiosis; high pollen and ovule sterility.
as-2 -- asynaptic-2
as2 Asynaptic meiosis; high pollen and ovule sterility.
as-3 -- asynaptic-3
as3 Asynaptic meiosis; high pollen and ovule sterility.
as-4 -- asynaptic-4
as4 Asynaptic meiosis; high pollen and ovule sterility.
as-5 -- asynaptic
as5 asynaptic meiosis; high pollen and ovule sterility.
as-6 -- asynaptic-6
as6 Delayed and asynaptic meiosis; chromatin degeneration; possibly two loci involve
Asc -- Alternaria stem canker resistance

Resistance to Alternia stem canker
at -- apricot

Yellow-pink color of fruit flesh
atn -- attenuata
at Narrow, pointed leaves, yellow-green, purple below; rigid cylindrical habit; thin shoots.
atv -- atroviolacium

Excess anthocyanin on leaves, stems, and fruits.
au 6 aurea
yg^6, yg-6, au^yg-6, yo Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
au (1k) aurea Knowles

au -- aurea

Bright yellow cotyledons and leaves; seedling etiolates; plant vigorous.
au tl aurea Torrey Lyons'
Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
au 3 aurea
au-2, au2 Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au 2 aurea
au^3 Elongated hypocotyl; yellowish foliage; smaller, slow-growing plant, dying prematurely.
au (2k) aurea Knowles second

au (1s) aurea Stubbe's au^2, au, brac Bright yellow cotyledons and leaves; some times whitish; corolla pale yellow; moderately vigorous in condine red; seedling etiolates; plant vigorous.
au w aurea
w616 Bright yellow foliage, often whitish; pale yellow corolla and stigma. Very weak.
auc -- aucta

Few branches; leaves broad, rounded, later becoming blistered, dark green; number of flowers and fruits reduced.
aur -- aurantiaca
aur1 Small, pointed, yellowish light-green pinnae; orange fruit.
aus -- austera

Plants always 1/3 to 2/3 normal size; pinnae finely divided, in some cases bent upwards; foliage darker and duller than normal.
aut -- aureata

Vireescent, flush of gold near growing point in early development, not detectable in older plants.
auv -- aureate virescent

Seedling 'gold dust' virescence; leaf segments pointed, sometimes fimbriate. Mature plant nearly normal.
avi -- albovirens
avi1 Smaller plant; foliage variously variegated yellow, white, grey-green; reduced fruit set.
aw prov4 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw prov5 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw prov3 without anthocyanin
aw Free of anthocyanin, same phenotype as a.
aw -- without anthocyanin
aba, ab, a179 Free of anthocyanin, same phenotype as a.
aw (1k) without anthocyanin
aw^2 Completely free of anthocynanin. Radiation induced in Money Maker.
aw-2 -- without anthocyanin-2

No anthocyanin at any stages; reduced growth rate, dense bushy habit.
B -- Beta-carotene

High B-carotene, low lycopene in ripe fruit.
B og Beta-carotene old gold og Corolla tawny orange; increased fruit lycopene.
B m Beta-carotene minutum
High B-carotene, low lycopene in ripe fruit. .
B c Beta-carotene crimson og^c,Crn,Cr,crn-2,cr-2 Increased fruit lycopene content; phenotype similar to B^og.
bc -- bicolor
bi Young leaves mottled brownish olive normal, later glazed dark green; short internodes; heterozygote intermediate for some characters.
Bco -- Brilliant corolla

Corolla pigment intensifier.
be -- blue-green

Small plant with blue-green leaves.
Blossom-end rot resistance

bg -- bursting resistance

Resistance of fruit to bursting.
bi -- bifurcate inflorescence

With f and j causes extreme fasciation of the "ring type".
bip -- bipinnata

Leaves highly divided, with smaller, acute segments.
bip prov2 bipinnata
bip Leaves highly divided, with smaller, acute segments.
bip-2 -- bipinnata-2
bip2 Slender, pointed, light gray-green cotyledons; doubly pinnate, rugose leaves.
bk -- beaked

Fruits with sharp points at stylar end.
bk 2 beaked

Fruits with sharp points at stylar end.
bk 3 beaked

Fruits with sharp points at stylar end.
Bk-2 -- Beaked-2

Sharp point at stylar end of fruit
bl -- blind

Stem terminating in first inflorescence; midribs may develop adventitious shoots.
bl 2 blind
to^2 Determinant growth; few flowered inflorescences; fasciated flowers; enlarged, leafy calxy; less extreme than bl.
bl to blind torosa to Determinate habit; firm, strongly uprolled pinnae; fasciated flowers and fruits.
bls prov2 baby lea syndrome
bls Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bls -- baby lea syndrome
alm Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bls prov3 baby lea syndrome
bls Anthocyaninless; restricted root system; short internodes, leaves, and trusses
bn -- blunt

Stamens absent, rarely present, with functional pollen, in greenhouse. Usually petaloid stamens adnate to pistil; leaf segments obtuse
bn sts blunt Stamen suppressor sts, sl-6 Stamens absent, rarely present, with functional pollen, in greenhouse; fewer blunt leaflets; with sp:+ 6-10 leafs between inflorescences.
Bnag-1 1 Beta-N-acetyl-D-glucosaminidase-1

br -- brachytic

Internodes shortened
brt -- bushy root

Radical branches early; radical and root profusely branched.
brt-2 -- bushy root-2

severely stunted growth (1/20) dense bushy growth of twisted roots.
bs -- brown seed

Brown seed color behaving as endosperm trait.
bs-2 -- brown seed-2

Brown seed color governed by genotype of endosperm.
bs-3 -- brown seed-3

Brown seed color determined by genotype of endosperm.
bs-4 -- brown seed-4

Brown seed color controlled by genotype of endosperm.
Bt -- bursting digenic inheritance
bg Susceptibility to fruit bursting.
btl -- brittle stem

Growth ceases with first flowering; stems brittle, with progressive defoliation; temperature-sensitive and expressed only under field conditions; B inefficient.
bu cin bushy compact inflorescence cin Internodes and inflorescences shortened; long petioles.
bu ab bushy abbreviata fru^ab Internodes and inflorescences shortened; long petioles.
bu cin-2 bushy compact inflorescence 2 cin-2 Internodes and inflorescences shortened; long petioles. Duplicate genes determining character similar to bu.
bu hem bushy hemiglobosa fru^hem Internodes and inflorescences shortened; long petioles. Phenotype like bu except leaves longer, more lax; heterozygote intermediate in some traits.
bu -- bushy
fru Internodes and inflorescences shortened; long petioles.
bul -- bullata

Leaf surface blistered and chlorotic with a network of darker veins; short internodes.
buo -- bullosa
buo1 Plants and seedlings larger in all parts; broad and rugose pinnae.
c prov3 potato leaf
c Number of leaf segments reduced
c prov4 potato leaf
c Number of leaf segments reduced
c prov6 potato leaf
c Number of leaf segments reduced
c prov2 potato leaf
c Number of leaf segments reduced
c prov5 potato leaf
c Number of leaf segments reduced
c -- potato leaf

Number of leaf segments reduced
c int potato leaf integerrima int Number of leaf segments reduced
car -- carinata

Shortened, keeled pinnae, dark gray-green color; deformed flowers; elongated fruit.
car-2 -- carinata-2
car2 Smaller plant; slender stems; smaller pinnae, nearly entire margin.
cb -- cabbage

Compact habit; leaves large and dark green; fewer locules.
cb-2 -- cabbage leaf-2

Larger plant parts; leaves epinastic and blistered.
constricting corolla

ccf -- cactiflora

Flower greatly fasciated; and roecium gynoecium and green tissues fused and distorted, enlarging into fruitlike structure; complete make and female sterility.
cd -- condensata

Seedling small; internodes shortened; leaf light grey-green with narrow, keeled segments; fruit round and relatively large.
ce -- cernua
ce1 Cotyledons lightly dotted in whitish yellow-green; lax, inclined plant.
Cf-? -- Cladosporium fulvum resistance

Resistance to specific races of Cladosporium fulvum
Cf-1 -- Cladosporium fulvum resistance-1
Cf, Cf1, Cfsc Resistance to race 1 and 3 of Cladosporium fulvum.
Cf-1 3 Cladosporium fulvum resistance-1
Cf-5, Cf5 Resistance to specific races of Cladosporium fulvum.
Cladosporium fulvum resistance-10

Cladosporium fulvum resistance-12

Cladosporium fulvum resistance-19

Cf-2 -- Cladosporium fulvum resistance-2
Cf2, Cfp1 Resistance to races of Cladosporium fulvum.
Cf-3 -- Cladosporium fulvum resistance-3
Cf3, Cfp2 Resistance to specific races of Cladosporium fulvum
Cf-4 -- Cladosporium fulvum resistance-4
Cf-8, Cf4, Cf-1^2 Resistance to specific races of Cladosporium fulvum.
Cladosporium fulvum resistance-5

Cf-6 -- Cladosporium fulvum resistance-6

Resistance to specific races of Cladosporium fulvum.
Cf-7 -- Cladosporium fulvum resistance-7

Resistance to specific races of Cladosporium fulvum.
Cf-9 -- Cladosporium fulvum resistance-9

Resistance to specific races of Cladosporium fulvum.
cfa -- conferta
cfa1 Very small, compact bush; strongly shortened internodes, young leaves.
cg -- congesta
cg1 Small, dense bush; late growth retardation; short internodes; pointed primary leaves.
ch -- chartreuse

Greenish-yellow subnormal petals; reduced fertility.
chln -- chloronerva

Upper leaves smaller and chlorotic interveinally; strong graft response.
chrs -- chrysanthemum sterile

Flowers fascicled 6cm across; many small pale yellow petals; stamens dialytic and without pollen; fruits deformed, orange-red fruits with brown sectors; poor fertility.
ci -- cincta
ci1 Small, irregular bush; many side shoots of equal length from lower leaf axils.
compact inflorescence-2

cir -- circumsaepta
cir1 Rounded, light gray-green pinnae, downrolled margin.
cit -- citriformis

In pm stock; Plant reaches 1/3 - 2/5 typical size; half erect, weakly branched; foliage color grey-green; terminal segment of primary leaves long and narrow; older leaves serrated; lemon shaped fruit with conspicuous stylar scar.
cjf -- confunctiflora

Flowers joined in pairs or triplets, otherwise malformed.
ck -- corky fruit

Pericarp cracked, exposed tissues becoming brown and leathery.
Cl -- Cleistogamous
Cl1 Flowers fail to open.
cl-2 -- cleistogamous-2
cl2 Flowers open only slightly; reduced fertility.
cla -- clara

Yellowish-green leaf color, purple veins and petioles; incompletely dominant.
clau ics clausa incisifolia ics Pinnae margins deeply incised, dentations acute.
clau prov2 clausa
clau Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau ff clausa

Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau -- clausa
ff, vc Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
clau vc clausa

Partly cleistogamous; leaves rugose, excessively divided; leaf lamina foreshortened, long petioled, flattened fruits.
cle -- collecta

Reduced growth rate; internodes shortened; small compact bush habit; leaf segments large, rounded, foliolules fewer.
cls -- clarescens

Foliage light yellow green early, becoming whitish green and blistered later, brilliant yellow green virescent; plants 1/4 typical size; growth very weak in field, setting fruits only in greenhouse.
clt -- coalita

Size and habit typically spl; primary leaves with entire end segments and only tiny laterals; mature leaves typically 5-segmented through fusion with end segment; segments always entire; resembles but is not allelic to sf.
clt-2 -- coalita-2

Terminal pinnae large, second order pinnae few and masked by 1st order pinnae, little serration.
cm -- curly mottled

Strong virus-like mottling and distortion of leaves and abortion of flowers; highly sensitive to environment.
cma -- commutata

dwarf with bright green foliage
Cmm -- Clavibacter michiganensis resistance

Resistance to Clavibacter michiganensis.
Cmr -- Cucumber mosaic resistance

Resistance to isolate 113A of cucumber mosaic virus, from L. chilense.
cn -- cana
ca Cotelydons and leaves gray-green especially on undersides; tiny and unbranched plants.
cna -- citrina

Foliage weak grey-green; petals light yellow; anthers greenish yellow.
co -- cochlearis

Reduced gray-green leaves with concave spoon-shaped leaflets; small plants of normal habit.
co-2 -- cochlearis-2
co2 Small leaves; deeply crenate, initially gray-green pinnae.
coa -- corrotundata
coa1 Plant small; pinnae fewer, broad, rounded, darker; flower parts short and broad.
com in complicata indiga in Small, dainty, nearly unbranched plants; shortened light gray-green leaves.
com -- complicata

Smaller, weakly branched erect plant; firm and keeled pinnae.
com-2 -- complicata-2
com2 Very small plant; short leaves, internodes; few side shoots; purplish shoot tips.
con -- convalescens

Yellow-green, paler at growing point; variably depressed growth.
con-2 -- convalescens-2
con2 Narrow cotyledons, dull light gray-green foliage, weakly yellow-green in growth zones.
cor -- coriacea

Smaller, unbranched plant; all parts reduced; short leaves with plicate segments.
Corynebacterium resistance-1

Corynebacterium resistance-2

cp -- compacta
cp1 Short leaves, smaller, dark green pinnae.
cpa -- composita
cpa1 Large, strongly branched, leafy inflorescence; jointless pedicel.
cpf -- cupuliformis

Cotyledons fused, cup-shaped, becoming coarse and enlarged later; transition to mature leaves without primary leaves; plants die at 3-4 leaf stage; line maintained via heterozygotes.
cpf-2 -- cupuliformis-2

Phenotype similar to cpf.
cpf-3 -- cupuliformis-3

Cotyledons adnate to form a cup; no primary leaves develop; maintained via heterozygotes.
cps -- compactilis

Plant smaller at all stages; leaves somewhat wider, lightly blistered and darker green.
cpt -- compact

Highly branched, compact plant; lax stems; pale, smaller leaves; reduced fruit set.
cr -- radial crack resistance

Resistance of fruit to radial cracking.
Cri -- Crispa

Very small in all parts; poorly viable without grafting; leaves light olive green, necrotic. Cri/+ necrotic spots on pinnae centers.
Crk -- Crinkled

Leaves like those of Cu but less condensed and segments dentate; homozygote lethal.
crt -- cottony-root

Overgrown root hairs with cottony appearance
cru -- corrupta
cru1 Smaller plant, irregular, sometimes distorted growth; spreading necrosis; firm, leathery, involuted brittle pinnae.
cs-2 -- corollaless-2

Apetalous; dialytic anthers; exserted stigma; self fertile, rarely sets fruits naturally.
cta -- contaminata
cta1 Smaller, flat, broad bush; many side shoots; at end of season stems and leaf midribs turn necrotic and brittle; also reduced fertility.
cta-2 -- contaminata-2
cta2 Leaves 1/4 normal size; stems with corky surface, variable necrosis of vascular tissue.
ctb -- cotyledon tops bifurcation

Bifurcation of cotyledon tops by a central vein.
ctr -- citrate concentration

Low citric acid in fruit (cv. Tondo Liscio) vs. high concentration, ctr (PI 263713 = L. esculentum).
ctt -- contracta

All shoots somewhat shortened, resulting in round bush habit; pinnae weakly serrated, plicate, and flattened; foliage somewhat lighter, and growing points are somewhat darker.
ctt-2 -- contracta-2

Plant half normal size at flowering; internodes shortened; leaves are narrow, pointed and blistered.
Cu -- Curl

Mid- and lateral veins and petiole greatly foreshortened producing crumpled leaf; homozygous viable.
cu-2 -- curl-2
cu2 Small, downcurled leaves; resembling Cu.
cu-3 -- curl-3

Dwarf, short and thick hypocotyls, curled cotyledons, and dense curly leaves; reduced fertility.
cua -- curta

Seedling and mature plant smaller, round bush habit; internodes and leaves shortened; leaves rounded and wider.
cul -- culcitula

Seedlings dark green with strong anthocyanin, later foliage dark grey-green; plants reach 1/3 to 2/5 typical size; internodes very short, habit cushion-like.
cup -- cupida
cus Extremely susceptible to drought damage at all stages; leaves small, yellowish, wilting, and eventually necrotic.
cur -- curvifolia

Leaves twisted, yellow-green; short internodes.
cv -- curvata
cu Young stem curves from node to node; dark rugose leaves; dwarf compact habit; heterozygote intermediate for some traits.
cv 2 curvata
acu Small plant; leaves, pinnae briefly stalked; leaf surface furrowed; older leaves strongly bent downward.
cv-2 -- curvata-2
cv2 Twisted cotyledons; short, rounded leaves; rugose, yellow-green pinnae.
cva -- conversa

Small, dense plant; variable fading of foliage.
cva-2 -- conversa-2
cva2 Small, compact plant, leaves, pinnae.
cvl -- convoluta
cvl1 Small, dense plant; short internodes; pinnae of oldest leaves involuted.
Cvx -- Convexa

Seedling and mature plant with narrow convex leaves, deeply veined and tending to roll downward on margins.
cy -- cyathiformis

Cotyledons united at one or both edges; lateral shoots suppressed; pinnae margins nearly entire.
d -- dwarf
rob^imm All parts foreshortened; leaves dark and rugose.
d x dwarf extreme
Extreme dwarf, all parts foreshortened; leaves dark and rugose; recessive to d and to +.
d provx-2 dwarf extreme-2 d^x All parts foreshortened; leaves dark and rugose.
d provcr-3 dwarf crispata d^cr All parts foreshortened; leaves dark and rugose.
d provcr-2 dwarf crispata d^cr All parts foreshortened; leaves dark and rugose.
d prov2 dwarf
d All parts foreshortened; leaves dark and rugose.
d im dwarf

All parts foreshortened; leaves dark and rugose.
d b dwarf broccoli
Extreme dwarf; more stunted than d^x; leaves reduced to small, dark green globs of curled, rugose tissue; flowers reduced, with tiny corolla segments.
d cr dwarf crispata rob^crisp Allele of phenotype intermediate between d and d^x.
d-2 -- dwarf-2
rob2, rob II, d2 Shortened internodes and leaves; thick, rugose, bluish gray-green leaves; not allelic with the mimic,d1.
d-3 -- dwarf-3
d3 Smaller, sturdy, erect plant; all parts foreshortened; rugose and dark-green leaves.
d-4 -- dwarf-4
d4 Mildly dwarf type; broad, shortened, crowded, darker pinnae; darker flower color.
d-5 -- dwarf-5
d5, d2, M-5 Slow growing plant with normal stem and leaf proportions; semi-sterile; classification good when three weeks old.
d-6 -- dwarf-6
d6, d5 Compact and small plant; rugose, and keeled pinnae, upper surface shiny dark green.
d-7 -- dwarf-7

Plants reduced to 10-12 inches tall; short internodes; leaves highly divided with strong anthocyanin.
d-8 -- dwarf-8

Plants reduced to 6-8 inches tall, short internodes; normal flowers but no fruit set in field.
dc -- decomposita
dc1 Slender, strongly sinuous and crenate pinnae; more secondary pinnae; micromutation.
dd -- double dwarf
d^xx Extremely retarded and highly modified dwarf.
de -- declinata

Great reduction in size of all parts; leaves dark yellow-green, leaflets roll toward underside.
deb -- debilis

Leaves emerge yellowish; necrotic mottling, whitish, and later brown, often constricting leaves in middle.
dec -- decumbens

Lax and decumbent habit; early fruiting.
dec-2 -- decumbens-2
dec2 After start of blooming plant becomes decumbent.
def -- deformis

Seedlings normal, later leaves becoming progressively reduced, filiform, and with adnate segments; deformed flowers and reduced fertility.
def 2 deformis
vit Leaves become progressively more deformed with twisted filiform leaflets.
def-2 -- deformis

Like def; leaves deformed after flowering.
Del -- Delta

Reddish-orange mature fruit color, due to inhibition of lycopene, and increase of delta-carotene.
deli -- deliquescens

Early habit dense later loosely parting; shortened yellowish leaves with narrow acute segments.
deli-2 -- deliquescens-2
deli2 Smaller compact bush; short internodes and leaves; all foliage is light yellow-green.
den -- densa

Typical spl seedlings; plant 3/4 normal size; stems and petioles strong; internodes somewhat shortened; leaf segments large and coarse; foliolules blistered; foliage color somewhat lighter.
dep -- deprimata

Slightly depressed growth; dainty, darker leaves.
depa -- depauperata

Very small plants with short internodes; leaflets bowed, color variable, and mostly yellowish.
det -- detrimentosa

Small, weakly branched and lax plant; yellowish foliage.
det 2 detrimentosa

Small, plant; dainty, pale yellow-green leaves; slender pinnae; less discoloration and premature dying than det.
det-2 -- detrimentosa-2

Plants reach 1/4-1/3 normal size; cotelydons narrow and tapered; leaves are shortened, segments small, keeled, margins serrated, and dull dark grey-green.
Df -- Defoliator

Homozygous seedlings retarded in growth, dark green, and die early; heterozygous are viable, but leaves become necrotic and fall prematurely.
dg -- dark green

Immature fruit color normal; darker green color appears as fruit develops, then persists until onset of ripening, high chlorophyll compared to normal or hp.
dgt -- diageotropica
lz-3 Plant habit prostrate due to reduced gravitropic response; growth retarded; stems and leaves droopy; cotelydons concave. Roots grow horizontally rather than downwards.
dgt dp diageotropica drooping leaf dp Leaves drooping, elongate, dark green; stems weak, slender, and prostrate.


Dia-2 1 Diaphorase-2

Dia-3 1 Diaphorase-3

Dia-3 2 Diaphorase-3


dil -- diluta

Leaves dull light green, small, and roughened; plants half normal size.
dil-2 -- diluta-2
dil2 Light colored, denser plant; short leaves, smaller pinnae.
dim -- diminuta

Somewhat smaller plants and leaves, shortened internodes; older leaves gray green with violet veins.
dim-2 -- diminuta-2
dim2 Very small plant, leaves, pinnae; much anthocyanin in hypocotyl, growth zones.
dis -- discolor

Leaf color light green with virescent growing regions, also with darker veins.
dis-2 -- discolor-2
dis2 Cotyledons first yellow-green, then whitish light-green; small compact plant.
div -- divaricata

Small squarrose plant with intercostally yellowish leaves and ventrally purple.
dl -- dialytic

Stamens not united; large trichomes partially suppressed and distorted. .
dl S dialytic Solanum
Anthers free, but trichome modifications of original dl absent; dominant to wild type.
dlb -- dilabens
dlb1 Small, compact plant with shortened yellowish leaves.
dlu -- dilucida

Plants reach 1/8-1/6 normal size, dwarf bush habit; foliage bright light green, light yellow-green virescent; petioles, stems, veins light violet; very low fruit set.
dm -- dwarf modifier
d2 Causes extreme dwarfing.
dmd -- dimidiata

Hypocotyl is strong; cotelydons are almost oval and arched downwards; internodes short and result in is small plants; leaf segments broad, coarse, and flat with 1 - 2 pairs of laterals grey-green; few foliolules develop later; corolla segments broad and flat; fruits few and seedless; maintained via heterozygotes.
dmt -- diminutiva

Shortened plant height and length of stem internodes, leaves, and pinnae.
dpa -- deparca

Small seedling and mature plant; little branching; leaf lighter at base, smaller, dark green; strong anthocyanin.
dpi -- densipinnata

Plant 1/2-3/4 normal size; small bush habit; cotyledons narrow with tapered tip; leaves dull grey-green; leaf segments large, crowded and fewer foliolules.
dpr -- depravata

Cotelydons and primary leaves strongly curled; internodes and leaves shortened; young leaves very asymmetrical, later deformed with long petiole.
dps -- diospyros

Fruit tissue is dusky orange.
dpy prov3 dumpy
dpy Leaves like those of d:x but internodes longer.
dpy prov2 dumpy
dpy Leaves like those of d:x but internodes longer.
dpy -- dumpy

Leaves like those of d:x but internodes longer; Leaves greatly condensed, rugose, dark green; internodes somewhat foreshortened
drt -- dwarf root

Reduced root system
ds -- dwarf sterile

Stunted plants with short internodes and reduced leaves; partially male-sterile.
dt -- dilatata
dt1 Plant small; pinnae yellowish; veins darker with uprolled margins.
dtt -- detorta

Cotyledons epinastic, irregularly crooked, many seedlings with 3 cotyledons; plants 2/5-3/5 normal size, less erect than normal; stems crooked and leaves and pinnae disordered; pinnae partly plicate, partly spiraled.
du -- dupla

Young plants typically splendens, later reach 2/3 size; habit half erect; branching reduced, particularly in the side shoot below the first inflorescence; leaf segments broad and doubled in number with many foliolules; strong anthocyanin in shoots exposed
dv -- dwarf virescent

Pale green virescent; plants always stunted.
dx-2 -- extreme dwarf-2

Resembles d:x in most respects; all parts extremely shortened; plant erect with no branching, attains half normal size; leaves very rugose and bright dark green above; flowers are small.
e -- entire
b Fewer fused leaf segments; midvein distorted
e 2 entire

Fewer fused leaf segments and distorted midvein, entire or broad leaflets
e prov3 entire
e Fewer leaf segments; midvein distorted
ec -- exserted carpels

Fruits vary from none to many exserted carpels.
eca -- echinata

Early growth typically pm and later reaching 2/3-3/4 typical size; from fruit set on main stem and branches grow vertically with dense to loose bush habit, lending porcupine-like appearance.
el -- elongated
e Elongated fruits as in Oxheart.
ela -- elata

Plants reach 3/4 normal size; dense, erect, and bush habit; internodes shortened; leaf segments large, wide, and strongly blistered.
ele -- elegans

Leaves reduced, yellow-green and dainty; plants tiny
ele 2 elegans
ang Reduced, sometimes chlorotic, narrow leaves; slow growth.
ele-2 -- elegans-2
ele2 Narrow, rounded cotyledons, margin rolled up; smaller, light yellow-green pinnae.
elo -- elongata
elo1 Elongated, weakly branched stems, initially erect, become procumbent.
elu -- eluta

Yellow green leaves; strong branching tendency; slight growth depression.
elu-2 -- eluta-2
elu2 Yellow-green to light gray-green leaves; late in season strong anthocyanin in shoots, stems, and veins.
em -- emortua
em1 Small plant; reduced side branching; spreading necrosis of older leaves leading to their premature death.
en -- ensiform

Sword-shaped sepals.
ent -- entire leaflet

Leaflets with entire margin.
ep -- easy peeling

Epidermis can be peeled from fruit without pretreatment.
Epi -- Epinastic

Severe epinasty of leaves, swelling of stems and petioles, prolific branching of roots. Partially dominant.
er -- erecta

Dwarf bushy plants with short internodes; shortened dark green leaves; heterozygotes intermediate.
er12.1 -- earliness

er-2 -- erecta-2

Plant vigorous, branches erect, internodes slightly shortened; leaves shorter, segments smaller, keeled and blue-green at tips; many flowered.
era -- eramosa
era1 Smaller, weakly branched plant; whitish zones in primary and later in leaves causing irregular pinnae development.
Erl -- Erectoid leaf

Leaves held at acute angle on stem; partially dominant.
Est-1 n Esterase-1

Est-1 5 Esterase-1

Est-1 4 Esterase-1

Est-1 3 Esterase-1

Est-1 1 Esterase-1

Est-1 2 Esterase-1

Est-2 1 Esterase-2

Est-3 1 Esterase-3

Est-4 5 Esterase-4

Est-4 7 Esterase-4

Est-4 6 Esterase-4

Est-4 8 Esterase-4

Est-4 3 Esterase-4

Est-4 1 Esterase-4

Est-4 2 Esterase-4

Est-4 4 Esterase-4

Est-5 1 Esterase-5

Est-6 1 Esterase-6

Est-7 1 Esterase-7

Est-7 2 Esterase-7

Est-8 1 Esterase-8

ete -- extenuata
ete1 Slender shoots; slender and thick pinnae; irregular growth; poor seed set.
eu -- eugenol concentration

High eugenol concentration in fruit.
ex -- exserted stigma

Exserted stigmas, or styles are twisted within anther tubes.
exa -- expassa
exa1 Branches partially decumbent; yellowish gray green older leaves turning yellow prematurely.
exl -- exilis
ex Very small plant; tiny plicate leaves of light gray-green color.
exs -- excedens
exs1 Smaller bush; short internodes, young leaves; older leaves leathery, and shiny.
f D fasciated fruit Dominant
Fruits many-loculed; phenotype more extreme than original allele of f; carpels unfused; dominant allele.
f -- fasciated fruit

Fruits many-loculed as in cv. Ponderosa.
fa -- falsiflora
fa1 Giant, vegetative, and highly ramified inflorescence; completely sterile.
far -- farinosa
far1 Smaller, weakly branched plant; many, deeply crenate, wavy, gray to yellow-green pinnae.
fc -- flaccescens

Internodes and leaves shortened; cotyledons narrow, wavy, light green; plants retarded at flowering, reach 1/5 normal size; leaf segments small, dark grey-green, tending to shrivel and die early.
fcf -- fucatifolia
fcf1 Small spreading plant; yellow or light green cotyledons, primary leaves, and lower leaves at later stages.
fd -- flecked dwarf

Retarded at all stages; leaves flecked with light green.
Fdh-1 1 Formate dehydrogenase-1

fe -- fertilis

Leaves roll dorsally; fasciated flowers, fruits.
Fenthion sensitivity

fer -- fe inefficient

Severe chlorosis beginning in first true leaves due to faulty iron transport in xylem.
Fgr H Fructose to glucose ratio hirsutum
Ratio of fructose to glucose in fruit approx. 1.5:1; partially dominant over L. esculentum allele
fgv -- fimbriate gold virescent

Leaf fimbriate margined, ephermerally "gold dust" virescent.
fi-2 -- filiforme-2

Phenotype like Schiemann's (1932), filiforme, equivalent to extreme wiry phenotype; cotyledons extremely narrow, grey-green; primary leaves also very narrow, later some leaves have only midveins, others with modified terminal segments and some epetiolate lateral segments; corolla with filiform segments; internodes very short, plant height averaging 20 cm.; no fruit set, line being maintained via heterozygotes.
fil -- filaris

Habit strongly branched and bushy; cotyledons slightly wavy; leaves wide, rounded, and often asymmetric with elongated petiole and filiform terminal segments; inflorescence very compound, petioles narrow, and fruit elongate.
fir -- firma

Broad blunt leaf segments; bushy cylindrical plants; foreshortened inflorescence.
fir-2 -- firma-2
fir2 Sturdy, compact seedlings; broad, rounded, rugose and dark green pinnae; small dark flowers.
fir-3 -- firma-3
fir3 Very small, compact bush; small, dark green, and rugose pinnae.
fis -- fistulosiflora

Corolla does not open although segments not fused.
fl -- fleshy calyx

Sepals fleshy, often curled.
fla -- flavescens

Leaves light with few segments; variably reduced plant size.
flav -- flavida

Leaves yellowish, mottled yellowish green, and small; heterozygote intermediate for some traits.
flav-2 -- flavida-2
flav2 Smaller, compact bush; slender stems, many side shoots; leaves yellowish light green.
flc -- flacca

Small, erect, nearly unbranched plant, and later spreading; short, overwilting, yellowish leaves.
fld -- flaccida
fld1 Small, weakly branched and erect plant; darker and involuted wilty pinnae, dying prematurely.
fle -- flexifolia
fle1 Very small plant, leaves with strong anthocyanin, borne at acute angle to stem in young plant.
fli -- filiform inflorescence

Filiform outgrowths and cupped leaves in late or terminal inflorescences; narrow corolla.
fms -- female-sterile

Distorted, poorly developed, highly sterile gynoecium; normal pollen production.
fn -- finely-netted

All true leaves light green with slightly darker veins.
fp -- fruit pox

Many small dark green spots on immature fruit, rupture prior to ripening.
fr -- frugalis

Small compact bush; older leaves dark-green; brittle, shiny.
FORL resistance

fra -- fragosa
fra1 Smaller, very brittle bush; broad cotyledonsc pinnae; heterozygote recognizable.
frg -- fragilis
frg1 Smaller, erect, nearly unbranched plant; short, dull light green leaves, yellowing prematurely.
fri 1 far red light insensitive

Insensitive to far red light; hypocotyls elongated relative to wild type under far red light.
Frl -- FORL resistance
Fr1, Fr-1 Resistance to Fusarium oxysporum f. sp. radicis- lycopersici; incompletely dominant.
Frly -- Frilly
Frl Leaf with serrate undulate margins.
fro -- frondea

Seedlings and plant habit and size normal; inflorescence highly proliferated; pedicels jointless; 4-parted corollae; segregation deficient in mutant homozygotes.
Frs -- Frosty spot
Nec Hypertrophic pustules along leaf veins followed by chlorosis then necrosis.
frt -- fracta

Plants attain 1/4 normal size; habit upright, weakly branched; leaf margins later turned upward; foliage dark green, shoots brittle.
Fs -- Fruit stripe

Fruit with dark green radial stripes opposite locules
fsc -- fuscatinervis
dkv Leaves yellow-green; veins always darker green.
ft -- fruiting temperature

Fruits can set at 40F.
fu -- fusiformis

Early growth habit is bushy and spindle shaped; leaves shortened, curved upwards, shiny yellowish, and paler at growing point.
fua -- fucata
fua1 Smaller plant; young pinnae yellow green, darker veins; older foliage variably lighter colored.
fug -- fulgida
fug1 Small plant; foliage variably whitish or yellow-green, sometimes zonally marked.
ful -- fulgens

Bright yellow virescent, older leaves becoming greener; golden yellow in field; yellow unripe fruit color.
ful 2 fulgens
ful1^2 Foliage at all stages light-green to light yellow-green; stems and leaf mid-ribs sometimes purplish.
ful-2 -- fulgens-2
ful2 Small plant; short internodes; light-green foliage, yellow-green growth zones.
ful-3 -- fulgens-3

Stunted virescentseedlings, overall brilliant yellow; closely resembling ful; mature plant bright yellow green.
fus -- fulgescens

Seedlings normal; plants normal size; foliage light green; growing points lighter; resembles, but is not allelic to ful.
fv -- flaveola

Seedlings small; leaf color light grey-green, a flowering light yellow green virescent; older leaves yellowish.
Fw -- Furrowed

Deeply furrowed, dark green cotyledons; severely stunted plant; homozygous inviable.
fx -- flexa

Young plants typically normal; main stem bends at start but becomes erect later, lending the plant a prostrate habit, similar to reptans(re).
fy -- field yellow

Foliage yellow at all stages and under all conditions.
g -- grooved

Inhibits modifiers, permitting expression of r.
ga -- galbina
ga1 Foliage at all stages variably white-gray-yellow-normal green; environmentally sensitive.
ga-2 -- galbina-2
ga2 Pinnae dark to light gray-green, yellow base; stems at all stages with strong anthocyanin; plant smaller in some years.
gas -- gamosepala
gas1 Small plant; gray-to-light-green foliage; wavy, irregularly crenate and rugose pinnae; partly connate sepals.
gbl -- globula

Plants always greatly reduced; hypoctyls; extremely short internodes; strongly branched; pinnae grey-green, coarse, crooked, downward; only observed in greenhouse; no flowers developed; maintained via heterozygotes.
Gdf -- Gold fleck

Small dark green spots on immature fruit, do not rupture but turn yellow on ripe fruit.
Ge c Gamete eliminator Condine Red
Most gametes abort when interacting with Ge:p.
Ge n Gamete eliminator neutral
neutral allele found in most genotypes
Ge p Gamete eliminator Pearson
Most gametes abort while interacting with Ge:p.
gf -- green flesh

Persistent chlorophyll giving ripe fruit purplish-brown color.
gfl -- globular flower

Corolla tips remain attached, resulting in globular form.
gfo -- grandifolia

Habit flatter; cotyledons and primary leaves variable whitish; leaves and segments large and light grey-green.
gh -- ghost
ab Incomplete chlorophyll deficiency, starting green, later breaking to white.
gh-2 -- ghost-2

Cotelydons yellowish; most leaves variegated.
gi -- gibberosa

Seedling relatively coarse; plant size normal; main stem semi-prostrate with reduced branching; leaves. mostly with 2 pairs lateral segments, very few foliolules, segments almost as broad as long, some terminal segments fused with laterals; less hairy stems and petioles.
gib-1 -- gibberellin deficient-1

Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-2 -- gibberellin deficient-2

Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-3 -- gibberellin-deficient-3

Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gib-3 x gibberellin-deficient-3

Gibberellin deficient; greatly reduced germination, and dwarf growth habit; leafs dark green, thick, and wrinkled; application of exogenous GA restores normal phenotype.
gil -- gilva

Yellow cotyledons; highly variable ratios.
gl -- glauca

Leaf segments narrow, acute yellow-green; virescent.
gl-2 -- glauca-2
gl2 Short internodes; small, dull, light gray-green leaves.
gla -- glaucicolor

Flat dense bush habit, strongly sidebranched; internodes shortened; foliage dull light grey-green; shoot tips with much anthocyanin; flowers small.
glau -- glaucescens

Leaves shortened, dull green to yellowish gray-green; small upright habit.
glau-2 -- glaucescens-2
glau2 Smaller plant; leaves dull gray-green; yellow-green growth zones.
glb -- globularis

Internodes and leaves shortened; leaves densely pinnate; yellow tinged.
glb-2 -- globularis-2
glb2 Dense, rounded bush; very short internodes; many side shoots; poor fruit set.
glc -- glaucophylla

Small plant; leaves light yellow or gray green, shortened.
glc-2 -- glaucophylla-2
glc2 Some pinnae larger, broader; pinnae light-green with lighter flecks.
glc-3 -- glaucophylla-3
glc3 Small plant and leaves of dull light grey-green.
glf -- globiformis
glf1 Very small, irregular bush; all plant parts reduced; many side shoots; multibranched inflorescences; young heterozygotes recognizable.
glg -- galapagos light green

Leaves pale gray-green, darker veins; plant small
glm -- glomerata

Seedlings normal; young plant with stronger anthocyanin; plants 1/3-2/3 normal size; shoots become necrotic at fruiting, resulting in condensed habit; later some branches become normal.
glo -- globosa

Internodes short; leaves short, pale green; incompletely dominant
glo 2 globosa
inx, intro Leaves tend to roll toward upper side, dull gray-light green; shortened internodes; excessively branched.
glo-2 -- globosa-2
glo2 Very compact, rounded bush; many side shoots; short internodes; small flowers, fewer clusters.
glo-3 -- globosa-3
glo3 Full sized erect bush; half sized, slender pinnae.
glu -- glutinosa
glu1 Dark green, shiny fruit with sticky epidermis; poor germination.
gm -- gamosepalous

Calyx and corolla segments tending to be connivent.
Got-1 1 Glutamate oxaloacetate transaminase-1

Got-1 2 Glutamate oxaloacetate transaminase-1

Got-2 1 Glutamate oxaloacetate transaminase-2

Got-2 4 Glutamate oxaloacetate transaminase-2

Got-2 n Glutamate oxaloacetate transaminase-2

Got-2 2 Glutamate oxaloacetate transaminase-2

Got-2 3 Glutamate oxaloacetate transaminase-2

Got-3 1 Glutamate oxaloacetate transaminase-3

Got-3 2 Glutamate oxaloacetate transaminase-3

Got-3 3 Glutamate oxaloacetate transaminase-3

Got-3 n Glutamate oxaloacetate transaminase-3

Got-4 2 Glutamate oxaloacetate transaminase-4

Got-4 n Glutamate oxaloacetate transaminase-4

Got-4 1 Glutamate oxaloacetate transaminase-4

Gp -- Gamete promoter

Strongly increased fertilization of gametes carrying it.
gq -- grotesque

Irregular androecium; pistil variably twisted, elongated, ridged, or lobed.
Gr -- Green ripe
gr Resembles gf, except that center of fruit turns red.
gra -- gracilis

Very small, unbranched, delicate plants; much reduced leaves yellowing at margins.
gra-2 -- gracilis-2
gra2 Very small plant, pinnae, short internodes and leaves.
gra-3 -- gracilis-3
gra3 Plant very small, erect, nearly unbranched; short internodes and leaves; pinnae and seedlings smaller.
grc -- gracillama
grc1 Smaller, weakly branched, half-erect plant; yellow-green, slightly involuted pinnae.
grf -- grandifructa
grf1 Broad, lax habit; fewer, large, flattened fruit; heterozygote intermediate.
grl -- gracilenta
grl1 Smaller, erect, weakly branched plant; large pinnae, close together, irregularly yellow green, normal green veins.
grl-2 -- gracilenta-2

Slender, pointed, pale green cotyledons; primary leaves delayed; dainty foliage; yellowish shoot tips.
grn -- granulosa

Base of large, multicellular trichomes greatly enlarged; associated chlorophyll deficiency and lack of vigor may be pleiotropic.
gro -- grossa

Cotyledons often more than 2; cotyledons and primary leaf segments short and broad, segments fewer, light grey-green; plants 2/3 normal height, upright, weakly branched; fruit and seed set reduced.
grs -- grisea
grs1, gr1 Smaller, light yellow green plant; dark gray-green growth zones.
gs -- green stripe

Radical green stripes in epidermis of unripe fruit; golden in ripe fruit
gt -- gametophytic factor

Reduced effectivity of female gametes.
Gx -- Gibberellin-exserted

Gibberellin-stimulated exsertion of style.
h -- hairs absent
H Large trichomes absent except on hypocotyl and at growing point; incompletely dominant.
ha -- hastata
ha1 Slender, long, pointed cotyledons; short leaves, with 2 pairs of slender, pointed pinnae; small, flat, dense bush.
he -- heteroidea

Small plant; internodes shortened; leaflets somewhat narrowed; older leaves glossy, brittle, and light green.
he-2 -- heteroidea-2
he2 Lax, low bush; slender stems, short internodes; small, keeled pinnae.
Hero -- Heterodera rostochiensis resistance

Resistance to potato cyst nematode (Globodera rostochiensis).
hg -- heterogemma
hg1 Small, lax plant; first inflorescence abnormal, numerous fasciated buds, elongated, unbranched peduncle.
hi -- hilara

Small plant, proportionately reduced; irregularly crenate and rugose and light-dark green patterned pinnae.
hl prov4 hairless
hl All herbage hairless; stems brittle
hl prov3 hairless
hl All herbage hairless; stems brittle
hl prov5 hairless
hl All herbage hairless; stems brittle
hl prov6 hairless
hl All herbage hairless; stems brittle
hl -- hairless

Large trichomeless stem cells; stems brittle
hl 2 hairless
cal, cal1 Smaller, weakly pubescent plant; dark-green leaves. Curved upwards to involuted.
hl-2 -- hairless-2
hl^prov6 All herbage hairless; stems brittle
hp-1 -- high pigment-1
hp, hp1, hp2, bs, dr Chlorophyll, carotenoids, ascorbic acid content of fruit intensified.
hp-1 w high pigment-1 WB3
Similar to hp-1, but more extreme phenotype.
hp-2 -- high pigment-2
hp Similar to hp-1; enhances all pigments of green and mature red fruits; in presence of u^+ allele, immature fruit are completely dark green.
hp-2 j high pigment-2 Jones hp Similar to hp-2.
Hr -- Hirsute

Long hairs on upper leaf surfaces.
Hrt -- Hirtum

Increased density of large trichomes; incompletely dominant.
hs -- hairless style

Style lack hairs.
ht -- hastate

Primary and mature leaves elongate, more entire, fewer lobed; abortive and deformed flowers.
hy -- homogeneous yellow

All vegetative parts yellow in all stages.
I -- Immunity to Fusarium wilt race 0

Resistances to Fusarium oxysporum f. lycopersici.
I-2 -- Immunity to Fusarium wilt race 2

Resistance to Fusarium oxysporum f. lycopersici race 2.
I-3 -- Immunity to Fusarium race 3

Resistance to Fusarium oxysporum f. lycopersici race 3.
ic -- inclinata

Small bush; short internodes, leaves; older leaves epinastic, yellowish.
ic-2 -- inclinata-2
ic2 Many slender side shoots; short leaves; narrow, smaller pinnae.
ica -- icana

Plants reach 3/4 typical size; habit denser with grey-green aspect, revealing some whitish-green of leaf under-surfaces; segments separate with light areas in younger leaves, partly fused, dirty yellow-grey in later leaves.
icn -- incana

Small plant, slow growing; whitish seedling and mature leaves latter with prominent purpling.
id -- indehiscens

Large pinnae; connate sepals; fruit irregularly cracked.
ida -- inordinata

Seedling typical pm; habit erect, irregular bush with less branching; some leaves with 2 pr. lat. segments, others unsegmented, in later growth becoming normal; term segments almost cordiform; foliolules strongly blistered; foliage color brilliant dark green
Idh-1 1 Isocitrate dehydrogenase-1

ig -- ignava

Dwarf upright, little branched; leaves relatively large, light green.
ig-2 -- ignava-2
ig2 Small plant; smaller leaves and pinnae of lighter color; short internodes.
ig-3 -- ignava-3
ig3 Cotyledons small, narrow; lax, flat bush; sturdy shoots, yellowish tips in field; large pinnae.
im -- impatiens
im1 Small, lax plant; reduced side shoots; darker, blue-green foliage wilts in strong sun and wind.
im-2 -- impatiens-2
im2 Small, flat bush; leaves short, small rugose, necrotic at tips; drought sensition.
imb -- imbecilla

Plant weak with few branches; younger leaves yellow-green then when older turning light green.
imb-2 -- imbecilla-2
imb2 Light green cotyledons, dark veins; smaller, lax plant; short, yellowish to light green leaves.
imp dia impedita

Smaller, weak plant; leaves chlorotic at margins; plant smaller than imp:eg.
imp eg impedita

Smaller, weak plant; leaves chlorotic at margins, like imp;dia but plant larger.
in-2 -- indiga-2
in2 Narrow, gray-green cotyledons; smaller plant; short internodes.
in-3 -- indiga-3
in3 Small plant; smaller pinnae; reduced branching; inflorescences close together and numerous.
ina -- inflexa
ina1 Lax growth, becoming partially decumbent.
inc -- incurva

Leaf segments weakly curved, undulate margins; stems, leaf veins and inflorescence crooked.
inc-2 -- incurva-2
inc2 Smaller plant; short internodes and leaves; long, narrow, keeled and brownish-green pinnae.
inf -- informa

Tiny plant with very short internodes; leaves small, plicate, and strongly down-curved.
ini -- inquieta
ini1 Small, proportionately reduced, variable plant; young pinnae involuted, lighter colored; large trichomes absent.
ino -- involuta
ino1 Smaller, variable, cylindrical bush; short internodes; involuted pinnae.
inr -- integer
inr1 Primary leaves almost undivided; long petioles; broad, rugose pinnae; few secondary pinnae.
ins -- inconstans
ins1 Small, weakly branched plant; phenotype, including leaf size, variable from year to year.
inta -- integrifolia
inta1 Lax, high bush; short leaves; rounded, crowded pinnae.
inv -- invalida

Small, dainty plants; irregular chlorotic flecks, later leaves deformed.
inx-2 -- introflexa-2
inx2, intro2 Plants diminutive in all respects; keeled to rolled, wilty pinnae.
Ip -- Intense pigment

Dark pigmentation of the unripe fruit.
irr -- irregularis

Leaves shortened, irregularly veined, dark green, tinged yellow.
ita -- inquinata
ita1 Young leaves in midseason develop small necrotic flecks, later coalescing into large ones; variable expressivity.
Itm -- Intumescences

Development of tumorous outgrowths on leaves and stems; heterozygote intermediate.
j -- jointless
lf Jointless pedicels; proliferates inflorescence.
j-2 in jointless-2
j2^in Jointless pedicel, with incomplete gene action. Pedicel joints with normal form, but do not separate when fruits ripen. Referred to as the 'arthritic' gene.
j-2 -- jointless-2
j2 Jointless elongate pedicel; teratological calyx; proliferated inflorescence; elongate fruit.
Jau -- Jaundiced

Yellow-green seedlings and young plants; homozygous lethal
jug -- jugata

Fasciated stems and fruits; short internodes; inflorescence almost sterile.
jug 2 jugata
jug1^2 Fasciated stems, flowers, fruits, more extreme than jug.
l prov4 lutescent
l Yellowish unripe fruits; premature yellowing of the leaves.
l -- lutescent
g Yellowish unripe fruits; premature yellowing of the leaves.
l 2 lutescent
rub Yellowish unripe fruits; premature yellowing of the leaves.
l prov3 lutescent
l Yellowish unripe fruits; premature yellowing of the leaves.
l-2 -- lutescent-2
l-3, l2 Premature yellowing of older leaves; yellowish unripe fruit.
La -- Lanceolate

Simple, entire, and small elongated leaves; slender stems, excessively branched; small fruits; homozygous inviable.
lae -- laesa

Small bush; axillary sprouts later necrotic; primary leaves dainty.
lan -- languida

Plant size and habit typical pm; until fruiting, variably light green virescent, later typical pm.
lower leaves albino fruit not ripening

lap -- lamprochlora
lap1 Small, lax plant; short, shiny leaves; end pinnae curved upwards (not seen in 97L).
lat -- lata

Habit broad at first; growth much depressed in field.
lax -- laxus
lax1 Plant lax, weakly branched; slender shoots; smaller, slightly rounded pinnae.
Lc -- Locule number

Locule number is reduced; associated with o.
le -- lembiformis
le1 Prostrate, smaller plant, proportionately reduced; keeled or involuted yellowish pinnae, ventrally purplish.
lep -- leprosa
lep1 Smaller, proportionately reduced plant; rapidly spreading necrosis on all older pinnae.
lg -- light-green
lme Light green foliage.
lg-2 -- light green-2
lg2, li, pg392 Cotyledons light yellow; leaves pale green; classification sometimes difficult.
lg-3 -- light green-3
lg3, lt, pg483 Light green cotyledons, leaves; former fade to yellow, latter to pale-green.
lg-4 -- light green-4
lg4 Yellow-green cotyledons, light green leaves.
lg-5 -- light green-5
lg5, lm, fy, yt Foliage uniformly light green
li -- limbrata

Seedling development variably reduced; first leaves slightly deformed with terminal segments broader at tip, in early stages pinnae tend to be finer, more strongly plicate, margins curled upward, and yellow-green. These characters disappear in mature plant.
liv -- livida

Plant erect, near normal size; cotyledons narrow, light grey-green, white-spotted; leaf segments long, narrow, slightly notched, blunt, dull grey-green; shoot tips bluish with anthocyanin.
Ln G Lanata

resembles Wo:m but is more hairy; all 3 genotypes viable and distinguishable.
Ln -- Lanata

Resembles Wo:m but hairs concentrated on stem; all 3 genotypes viable and distinguishable.
lo -- locule number reduced

Reduced locule number.
loa -- low acid

Low titratable fruit acidity.
loc -- locule number increased

Locule number increased.
lop -- longipes
lop1 Small, somewhat spreading plant; broad, weak gray-green pinnae; long petioles; variable expression.
Lpg -- Lapageria

Leaves small, dark green, glossy; flowers campanulate; homozygote sterile.
ls -- lateral suppresser

Few or no axillary branches; corolla suppressed; partially male sterile
ls 2 lateral suppresser

Few or no axillary branches; corolla suppressed; partially male sterile.
lt -- laeta
lt1 Smaller, weakly branched plant; light gray-green foliage; growing points, later entire plant, yellow-green.
lt-2 -- laeta-2
lt2 Dainty, light green foliage; shoot tips with yellowish pinnae and in late season has bluish veins, petioles.
ltf -- latifolia

Cotyledons and pinnae extremely broad proportioned.
lu -- luteola

Light yellow corolla. .
luc -- lucida

Leaves bright green, yellow-green at growing point.
luc-2 -- lucida-2
luc2 Smaller, lax bush; short stems; small, light green leaves.
luc-3 -- lucida-3
luc3 Plant and leaves small, light green to light olive.
lur -- lurida
lur1 Yellow-green cotelydons and primary leaves; dull light green later foliage.
lut -- lutea

Leaves yellow-green, darker veins; virescent.
lut-2 -- lutea-2
lut2 Primary leaves flecked yellow-green; pinnae dull light green, yellowing early.
lut-3 -- lutea-3
lut3 Cotyledons yellowish light green, small plant and leaves of dull light green.
lux -- luxurians
lux1 Young plant larger, fewer side shoots; erect, long internodes.
Lv -- Leveillula taurica resistance

Resistance to Leveillula taurica powdery mildew.
Lx -- Lax

Elongate, acute-segmented, pendent leaves.
lyr -- lyrate

First leaves entire; later fan shaped; female-sterile.
lz -- lazy

Normal seedlings, 3 weeks later become strongly geotropic; weak branching.
lz-2 -- lazy-2

Plant habit strongly prostrate due to reversed geotropic response of shoots (roots normal).
m -- mottled

Leaves and cotelydons with tiny flecks of paler green.
m-2 -- mottled-2
m2, mo, md Many small chlorotic spots on leaves; plant stunted; temperature-dependent.
M-3 -- Mottled-3
M3, M345 Variable sectoring for chlorophyll deficiency; homozygous lethal.
ma -- macrocarpa

Leaves clearly longer; fruit large, smooth.
maa -- macra

Branching inhibited, erect habit; leaf segments and flowers reduced size.
mac -- maculata
mac1 Small plant; necrosis on upper or lower leaf surface spreading from margin.
mad -- marcida
mad1 Small plant; dark-green, slightly wilty foliage.
Mae-1 1 Malic enzyme-1

Allele of S. lycopersicoides, slightly retarded from '+'.
mal -- maculosa
mal1 Young plant leaves have irregular, yellowish light green flecks, disappear later.
mar -- marcescens

Small plant; leaves dark-green, lightly wilting.
marm -- marmorata

Marbling of several tones of white and light green on leaves, strongest expression in greenhouse.
marm 2 marmorata
marm1^2 Marbling of several tones of white and light green on leaves, strongest expression in greenhouse.
marm-2 -- marmorata-2

Cotyledons variably white, yellow, yellow-green or green with variegated true leaves.
mat -- matura
mat1 Lax, flat bush, retarded in growth; short leaves; half-sized pinnae.
mc -- macrocalyx

Sepal large and leaf-like; pedicel joint arthritic; inflorescence indeterminant.
mcn -- maculonecrotic

Seedling 2/3 normal size with virescence, yellow spots becoming necrotic, mature leaves often deformed, especially mid-region, also virescent at maturity.
mcr -- multicolor

Seedling typical spl; plant 3/4 size; habit spreading; foliage color yellow green early to dark midveins with necrotic spots later, finally with variable whitish-yellow-green aspect.
mcs -- macrosepala

Size and habit normal; inflorescence elongate, indeterminate; corolla enlarged; closed, but opening when touched; calyx with similar features; older leaves uncurled to reveal undersides.
Mdh-1 2 Malate dehydrogenase-1

Mdh-2 1 Malate dehydrogenase-2

Malate dehydrogenase-3

Mdh-4 1 Malate dehydrogenase-4

Me -- Mouse ears

Leaves 3-4 pinnately compound with clavate segments; shortened internodes; homozygous viable; incompletely dominant.
med -- mediocris
med1 Smaller plant, proportionately reduced.
mel -- melongenoida
mel1 Sometimes lax habit; oval fruit.
men -- mendosa

Seedlings variable size; plant reaches half normal size; leaves always deformed, segments reduced, curled, very blistered; flowers deformed, style exserted from bud stage.
mes -- methyl salicylate concentration

High levels of methyl salicylate in fruit of certain lives.
mf -- multiformis

Growth always retarded; thin branches; internodes and leaves shortened, segments small, oval, almost entire; very few foliolules, foliage starts grey-green, later dull green.
mgn -- marginal necrotic

Seedling 1/4 normal size; cotyledons and leaves yellowish, becoming necrotic first at tips and margins; mature plant subnormal in size and leaves with brown necrotic blotches.
Mi -- Meloidogyne incognita resistance

High level resistance to root-knot nematode.
Mi-3 -- Meloidogyne incognita resistance-3

Resistance to Mi-breaking strains of Meloidogyne incognita, including at high temperatures.
Mi-4 -- Meloidogyne incognita-4

Heat stable resistance to Meloidogyne incognita and M. javanica
mia -- minuscula

Seedlings small; nearly no branches, erect habit; leaf segments small, keeled, dull dark grey-green; inflorescence proportionately large.
mic -- microcarpa
mic1 Leaves, stems marked in gray-green; lighter yellow petals, uprolled margins; ridged or yellow-fissured fruits.
min -- minutula

Growth always retarded; short, weak branches; internodes and leaves very shortened; stems grey-blue; foliage light grey-green.
mlt -- malate concentration

Low malic acid levels in fruit versus high levels in mltD.
mn -- minuta
mi Small, upright and almost unbranched plants; shortened leaves prematurely yellowing.
mn-2 -- minuta-2
mn2 Very small plant, pinnae; short leaves.
mnt -- miniature

Small plant; reduced branching and fruit set.
mnx -- extreme miniature
min^ex Severely stunted chlorina mutant with 1cm internodes; flowers only after grafting on normal.
Mo(I) -- Modifier of I

Heterozygote overdominant.
Mo(I)-2 -- Modifier-2 of I

Exerts less effect of I than Mo(I).
mo-B -- modifier of B
mo(B), moB, I^B, i^B Modifier of B. Increases content of beta carotene in presence of B.
Moi -- Modifier inducer

Permits expression of Mo(I) and MO(I)-2 only in xx genotype.
mon -- monstrosa

Very tiny, upright, and unbranched plants; leaves much reduced, down-curved, and ball shaped; heterozygote intermediate for some traits.
mor -- morata
mor1 Smaller plant, reduced side branching; yellowish light green foliage.
mps -- miniature phosphorus syndrome
ph Very retarded and all parts extremely reduced; foliage chlorotic and strongly flushed with anthocyanin, resembles pds.
ms -- male sterile
ms-1, ms1 Pale, shrunken anthers; no pollen; hybrid stock
ms-02 -- male-sterile-2
ms2 Anthers pale, shrunken; no pollen
ms-03 -- male-sterile-3
ms3 Anthers variably small, discolored, occasionally fertile
ms-03 42 male-sterile-3
ms-42 Normal anthers, low amount of good pollen in some genetic backgrounds.
ms-04 -- male-sterile-4
ms4 Pale, shrunken anthers; a few aborted pollen grains.
ms-05 -- male-sterile-5
ms5 Abnormally small flowers; very pale and greatly shrunken anthers; usually no pollen.
ms-06 -- male-sterile-6
ms6 Shrunken, pale anthers; no pollen.
ms-07 -- male-sterile-7
ms7 Anthers near normal in size and color; pollen aborted in tetrads.
ms-08 -- male-sterile-8
ms8 Abnormally small flowers with exserted stigmas; plale shrunken anthers; no pollen
ms-09 -- male-sterile-9
ms9 Anthers nearly normal; no pollen
ms-10 -- male-sterile-10
ms10 Abnormally small flowers; small, very pale anthers; greatly exserted stigmas; no pollen.
ms-10 35 male-sterile-10
ms-35, ms35 Anthers small, slender, pale; empty PMCs in locules; breakdown in different stages of meiosis; stigmas exposed.
ms-10 36 male-sterile-10
ms-36 Anthers small, very pale; no pollen; flowers very small
ms-11 -- male-sterile-11
ms11 Very pale, shrunken anthers; aborted pollen, free or in tetrads.
ms-12 -- male-sterile-12
ms12 Abnoramally small flowers; nearly normal-colored but shrunken anthers; no pollen.
ms-13 -- male-sterile-13
ms13 Nearly normal anthers; free aborted pollen.
ms-14 -- male-sterile-14
ms14 Anthers shrunken; very pale; aborted pollen
ms-15 -- male-sterile-15
ms15 Anthers dwarfed, very pale; no pollen; flowers small; stigmas exserted
ms-15 26 male-sterile-15
ms26, ms-26 Abnormally small flowers with exserted pistils; very pale, dwarfed anthers; no pollen.
ms-15 47 male-sterile-15
ms-47 Anthers dwarfed, very pale; no pollen; flowers small; stigmas exserted
ms-16 -- male-sterile-16
ms16 Pale shrunken anthers; clumped aborted pollen.
ms-17 -- male-sterile-17
ms17 Pale shrunken anthers; no pollen.
ms-18 -- male-sterile-18
ms18 Exserted stigmas; slightly pale, very shrunken anthers; no pollen;
ms-19 -- male sterile-19
ms19 Slightly shrunken anthers; stigma mostly protruding.
ms-20 -- male sterile-20
ms20 Anther tips equal or exceed stigma.
ms-21 -- male sterile-21
ms21 Anthers and style of equal length; flowers slightly smaller.
ms-22 -- male sterile-22
ms22 Anthers normal length, shrunken; flower color slightly lighter.
ms-23 -- male-sterile-23
ms23 Anthers slender and paler; no pollen; meiosis delayed and prolonged; stigmas depressed.
ms-24 -- male-sterile-24
ms24 Anthers slightly more slender and paler; 100 percent aborted pollen; meiosis normal stigmas mostly exposed.
ms-25 -- male-sterile-25
ms25 Anthers slender, foreshortened, pale; stigmas mostly exposed.
ms-27 -- male-sterile-27
ms27 Anthers slender, diverging distally, pale; stigmas mostly not exposed.
ms-28 -- male-sterile-28
ms28 Anthers very slender, slightly shorter, pale; stigmas mostly exposed.
ms-29 -- male-sterile-29
ms29 Anthers slender, often separate below, pale, often greenish; stigmas mostly exposed.
ms-30 -- male-sterile-30
ms30 Stamens highly modified, very slender, free twisted, pale or greenish yellow; stigmas 100% exposed.
ms-31 -- male-sterile-31
ms31 Stamens smaller, very pale; stigmas 100% exposed.
ms-32 -- male-sterile-32
ms32 Stamens greatly reduced and shrunken, very pale, often brown.
ms-33 -- male-sterile-33
ms33 Anthers very irregular, yellow-green; no pollen; breakdown in early meiosis; stigmas exposed.
ms-34 -- male-sterile-34
ms34 Anthers short, pale; no pollen; meiosis normal, possibly delayed; stigmas exposed.
ms-36 -- male sterile-36
ms36 Anthers slightly shrunken and greenish.
ms-37 -- male sterile-37
ms37 Flowers appear normal, but have no pollen.
ms-38 -- male-sterile-38
ms38 Flowers small; anthers pale; no pollen; stigma not exposed
ms-38 40 male-sterile-38
ms-40 Flowers small; anthers pale (matching corolla); no pollen; stigma inserted.
ms-39 -- male-sterile-39

Flowers possibly more fasciated than normal; stamens pale and slender; no pollen; stigma variably exposed; plant somewhat chlorotic and weak.
ms-41 -- male sterile-41

Anthers slghtly smaller, paler than +; all pollen aborted.
ms-43 -- male sterile-43

Few or no stamens produced, pollen viable
ms-44 -- male-sterile-44

Solanifolium leaf shape; small flowers with protruding style and divided stigma.
ms-45 -- male-sterile-45

Anthers long, slender, and pale; stigmas usually not exposed; breakdown sporogenesis and vestgia resorbed in later anther development.
ms-46 -- male-sterile-46

Anthers long, slender, and pale; stigmas usually not exposed; breakdown sporogenesis and vestgia resorbed in later anther development.
Ms-48 -- Male-sterile-48

Anthers pale, slender; completely male sterile; only small, abortive cells visible, no pollen grains.
ms-49 -- male-sterile-49

Anthers pale, slender.
ms-50 -- male sterile-50

Anters pale, shrunken. High level of male-sterility; (but some pollen produced) anthers tend to brown and shrived distally.
Ms-51 -- Male-sterile-51

Dominant male-sterile mutant, similar to Ms-48.
mt -- midget

All parts of plant reduced; high sterility.
mta -- mutata
mta1 Small broad bush; internodes short; leaves long; light yellow-green, virescent in early stages.
mts -- mortalis
mts1 No inflorescences even if grafted on normal stock; smaller bush; strongly reduced internodes and leaves.
mu -- multinervis

Dainty young leaves, and light green intercostally.
mu 3 multinervis
rv-3 Seedlings with strong virescence of light green color between veins of normal green.
mua -- multifurcata
mua1 Small plant, all parts proportionately reduced; first inflorescence multibranched.
muf -- multifolia

Leaves elongate, rugose, highly divided, gray-green.
mult -- multiflora

Very greatly increased number of flowers per inflorescence.
mun -- multinata
mun1 Small bush; many equal side shoots; short internodes and leaves.
mup -- multiplicata
mup1 Enlarged, multibranched inflorescences; variable whorled flower parts; fruits fewer
mut -- mutabilia
mut1 Small plant, reduced branching; midseason leaf color weak gray-to-dark green.
muv -- multivalens
muv1 Plants and leaves small, light green, inflorescences close together
muv-2 -- multivalens-2
mus1 Small plant, shiny, shorter leaves; light yellowish growing points.
mux -- multiplex
mux1 Stems often end abruptly in leafy structure; inflorescences irregularly branched to fasciate; jointless pedicels swollen at calyx base; petaloid sepals.
n -- nipple-tip
nt Nipple tips at stylar end of fruits.
n-2 -- nipple tip-2

Nipple tips at stylar end of fruits.
n-3 -- nipple tip-3

Nipple tips at stylar end of fruits.
n-4 -- nipple tip-4

Nipple tips at stylar end of fruits.
na -- nana

Tiny plant, internodes and leaves very short, plicate and rugose.
na-2 -- nana-2
na2 Very small, erect bush; tiny, narrow, rugose pinnae, often with necrotic flecks.
nc -- narrow cotyledons

Narrow cotelydons and slow growth.
nd -- netted
m-4 Primary leaves attenuated and chloroticlater leaves less extreme; chlorophyll concentrated around veins of later leaves.
ndw -- necrotic dwarf

Semi-dwarf habit, epinastic leaves, necrosis in stems and leaves.
ne -- necrotic

Necrotic; with Cf:2 causes progressive necrosis of leaves.
ned -- necrodeformis

Leaf chlorosis, turns necrotic; some plants stunted and distorted; temperature dependent.
neg -- neglecta

Plants small, weakly branched; leaves pale, darker veined, becoming necrotic, and prematurely shed.
neg ne-2 neglecta necrosis 2 ne-2, ne2 Leaves with many grayish necrotic spots; plant small, weak, killed by defoliation.
neg-2 -- neglecta-2
neg2 Plants diminutive in all parts; pinnae become necrotic.
neg-3 -- neglecta-3
neg3 Smaller, weakly branched plant; dark gray-green pinnae, necrotic flecks at tips, dying prematurely.
nf -- no flowers

Scanty flower production.

ni-2 -- nitida-2
ni2 Narrow cotyledons, pinnae; older shoot tips blue-green and slender.
Nir-1 1 Nitrate reductase-1

nit -- nitidula
nit1 Lax, low bush; small, dark gray-green leaves, slightly glossy above.
nor -- non-ripening

Very retarded pigmentation, non-softening, and crack resistance of fruits.
nor 2 non-ripening

Very retarded pigmentation, non-softening, and crack resistance of fruits.
nor A non-ripening

not -- notabilis

Dainty and delicate yellow-green leaves; tends to wilt in dry or hot weather.
Nr -- Never ripe

Fruits ripen slowly to dirty orange color; homozygous viable.
Nr-2 -- Never ripe-2

Fruit flesh turns slowly to yellowish green, remains firm
nt -- nitens

Plants reach 1/3 normal size, branching reduced; internodes and leaves shortened; leaf dark green, upper surface bright.
nv -- netted virescent

Pale interveinal areas of cotyledons and true leaves; older leaves never with full green color.
o -- ovate

Pear shaped fruits.
o 1(2s) ovate
ol^2 Ovate fruits.
O. 1 Oval
ol Elongoate fruits with low locule number.
O. 1(1s) Oval
ol^3, ov, ov1, Small compact bush; short internodes; small leaves and pinnae; ovate fruits.
ob -- obscura

Smaller, weakly branched plant; early foliage dark-green.
ob-2 -- obscura-2
ob2 Smaller bush; short internodes; smaller, broad, compacted, dark-green pinnae.
obl -- oblate fruit

Oblate fruit shaped.
obv + obscuravenosa clear vein
Leaf veins appear clear with back lighting due to absence of chloroplasts in epidermal cells beneath veins.
obv prov1 obscuravenosa

obv prov2 obscuravenosa

obv -- obscuravenosa

Leaf veins appear green with back lighting, due to presence of chloroplasts in epidermal layer beneath veins.
oc -- ochroleuca

Leaves discolored and becoming white variegated; exposed unripe fruit is yellowish gray.
Od -- Odorless

Herbage with little or no characteristic tomato odor.
old gold

Ol-1 -- Oidium lycopersicum-1

Resistance to Oidium lycopersicum; incompletely dominant
Ol-3 -- Oidium lycopersicum resistance-3

Resistance to Oidium lycopersicum; incompletely dominant.
oli -- olivacea

Very small plant; small, malformed olive-green leaves, dying prematurely.
oli 2 olivacea
oli1^2 Small plant; pinnae more divided, short, variously light-olive-dark green, wavy margin, dying early.
op -- opaca

Lighter leaf color, yellow at growing point.
op-2 -- opaca-2
op2 Small plant with light green foliage, particularly in growth zones.
opa -- opacata
opa1 Smaller somewhat spreading plant; yellow-green foliage.
or -- ordinata

Plant size and habit typical pm; until fruiting, variably light green virescent, later typical pm.
Ora -- Orobanche aegyptica resistance

Resistance to broomrape based on single gene action.
os -- oligosperma
os1 Small, weakly branched, half erect plant; distorted cotyledons; dark green leaves, fewer pinnae.
ot -- obtusa
ot1 All leaves short, broad, dark green, weakly shiny above.
ova -- ovata

Seedlings smaller, branches slender, internodes shortened; foliage early whitish light green, later normal but light green virescent; fruits variably egg shaped.
ovi -- oviformis
ovi1 Broad, keeled, more crenate pinnae; long oval fruit.
p 2 peach
nor^3 Dull-colored fruit epidermis, from mature green to ripe. Phenotype similar to nor^2, but pericarp more pink-colored and skin less strong, possibly equivalent to nor.
p -- peach

Fruits with dull hariry surface.
pa -- parva
pa1 Very small plant; few side shoots; short internodes; small leaves and pinnae.
pa-2 -- parva-2
pa1, pa2 Small, nearly unbranched, erect plant; firm leaves, weakly curved upwards.
paf -- pauciflora
paf1 Fewer flowers per inflorescence; small plant and pinnae.
pai -- pseudoanthocyanin incomplete

Strong purple of hypocotyls fades shortly after germination to dusky green but returns to positive level by flowering time.
pal -- pallida

Light green color; paler corolla; plant size variable; foliage color dominant.
pap -- paupercula

Seedlings typical praematura, plants reaching 1/3-2/5 praematura size; branching variably reduced; susceptible to wilting from flowering on.
par -- parca
par1 Smaller, weakly branched, half erect plant; fewer boat-shaped to involuted pinnae, purplish ventrally.
pas -- pallescens
pas1 Small plant; dull light green foliage.
pas-2 -- pallescens-2
pas2 Small, narrow, light green, keeled pinnae.
pat -- parthenocarpic fruit

Parthenocarpic fruit; closely linked with sha (short anthers).
pat-2 -- parthenocarpic fruit-2

Parthenocarpic fruit.
pau -- pauper

Plants tiny, weak, and unbranched; tend to die before fruiting; marked reciprocal graft influence with normal.
pc -- precocious centromere division

Centromeres divide prematurely in meiosis starting in anaphase I and completed in all by prophase II; completely pollen sterile, highly egg sterile.
pca-2 -- proclinata-2
pca2 Smaller, dainty plant; short, light gray-green leaves; full-sized flowers.
Pch + Photoperiodic chlorosis

Chlorosis under constant illumination; apparently in all L. esculentum; dominant absence allele in other species.
pct -- polycot

Multiple cotyledons, smaller leaves with fewer segments, entire margins, grayish, strongly convex, deep veined. Stunted growth. Inflorescence large, compound, epinastic. Flowers fasciated. Corolla segments narrow.
pcv -- polychrome variegated

Fine striated variegation for white, green and various intensities of gray-green on distorted leaves.
pd -- pigment diluter

Reduces plant chlorophyll and fruit pigment, the latter by 50%.
pdc -- pudica

Seedling very retarded and dark colored; cotyledons and first true leaves plicate, arched and hooded; mature plant greatly retarded; leaves narrow, strongly plicate, and tips acute.
pds -- phosphorus deficiency syndrome
Ph-oid Development retarded; leaves reduced, distorted, blotched dull yellow green and flushed with anthocyanin.
pdw -- pale dwarf

Plant small; leaves pale yellow, not turning green.
pe 2 sticky peel
pe-2 Phenotype similar to pe (sticky fruit epidermis); waxy fruit.
pe -- sticky peel

Sticky fruit epidermis.
pen -- pendens

Broadly spreading, drooping, glossy, and yellowish pinnae.
per -- perviridis

Leaves darker green, dropping early, anthocyanin strong; very small, slow-growing plants.
pet -- penetrabile
pet-2, pet2 Plant small, stocky shoots; short internodes and leaves; rugose pinnae; heterozygote intermediate at season's end.
pf -- parviflora

Plants reach half normal size; internodes and leaves shortened; leaf segments small, narrow, keeled, dark grey-green; branch tips purplish; flowers smaller, petioles narrow and slightly paler.
pg -- pale green
pg329 Cotyledons first pale yellow, turn gray-green, which is mature plant color; slow growing.
pg-2 -- pale green-2
pg2, pa, pg390 Cotyledons and leaves pale-green.
pg-3 -- pale green-3
pg3, pl, pg391 Cotyledons pale gray-green; leaves fading to dull yellow color.
Pgdh-1 -- 6-Phosphogluconate dehydrogenase-1

Pgdh-2 1 6-Phosphogluconate dehydrogenase-2

Pgdh-3 1 6-Phosphogluconate dehydrogenase-3

Pgi-1 2 Phosphoglucoisomerase-1

Pgi-1 1 Phosphoglucoisomerase-1

Pgm-1 1 Phosphoglucomutase-1

Pgm-2 1 Phosphoglucomutase-2

Ph -- Phytophthora infestans resistance
PiT, TR1 Resistant to Phytophthora infestans race T:0, susceptible to race T:0.
Ph-2 -- Phytophthora infestans resistance

Phytophthora infestans resistance.
Ph-3 -- Phytophthora infestans resistance

Phf -- Phytophthora infestans field resistance

Incompletely dominant.
Phf-2 -- Phytophthora infestans field resistance2

Incompletely dominant.
pi -- pistillate

Flowers usually lack stamens and are otherwise modified.
pi-2 -- pistillate-2

Plant size and vigor reduced; inflorescences tending to abort; flowers small with only calyx and pistil; calyx subnormal, tightly adpressed to pistil; female fertility and/or fruit set affected.
pic -- picta

Leaves at first normal green, later turning yellowish then brown necrotic above, some eventually brown below.
pic-2 -- picta-2
pic2 Young pinnae with marginal necrotic flecking, later more numerous.
pic-3 -- picta-3
pic3 Plant diminutive in all parts; pinnae light yellow-green turning necrotic at tips, dying early.
pic-4 -- picta-4
pic4 Scattered necrotic flecks on smaller, light gray to olive green pinnae.
pin -- pinnosa
pin1 Small, flat bush; short internodes, leaves; keeled, sometimes yellowish, light green pinnae.
pl -- perlucida
pl1 Light green, narrow pinnae, yellowing prematurely.
pla -- plana

Small plant, stiff when young, yellowish to light green leaves darker veins.
pla-2 -- plana-2
pla2 Longer hypocotyl; larger cotyledons and pinnae.
pli -- plicata

All parts small; leaves dark, yellowish, plicate; strong anthocyanin.
pli-2 -- plicata-2
pli2 Small, lax bush; slender shoots, short internodes; smaller gray-green pinnae.
pm -- praematura
pm1 Small plant; dainty, shorter, sometimes yellowish leaves; fruit ripen early.
pm-2 -- praematura-2
pm2 Short internodes; small, rounded, crowded, dark green pinnae; fruit ripen early.
pma -- praemortua

Plants reach 1/4 normal size, weakly branched; leaves reduced; starting at flowering: stem epidermis becomes dry; brown, silvery, leaves gradually necrose; fruit set good for plant size.
Pn -- Punctate

Base of trichomes accumulate anthocynanins.
pol -- polylopha

Short internodes; leaves plicate and twisted; fruit flat and fasciated.
pol-2 -- polyopha-2
pol2 Main, lateral shoots end in large branched inflorescence with large, fasciated central flower; fruit flattened 4 to 9 locules.
Pox -- Poxed fruit

pp -- polyphylla
pp1 Longer, more divided, lighter-colored leaves, micromutation.
ppa -- purpurea

In early stages indistinguishable from elu; as fruits set, becomes distinguishable by its high anthocyanin content.
ppa 2 purpurea

In early stages indistinguishable from elu; as fruits set, becomes distinguishable by its high anthocyanin content.
ppi -- paucipinnata

Plants always retarded, weakly branched, shortened internodes; leaves often with 1 pair of segments, acuminate, slightly keeled; reduced fruit set.
pr -- propeller

Cotyledons propeller like; true leaves greatly modified.
pra -- praeclusa

Cotyledons short, broad, bent downwards on strong hypocotyls; plants(seen only in greenhouse) of variable height 10-40 cm; habit upright with no branches; leaves extremely foreshortened and crupled with 1-2 partial segments, dark green, strongly blistered and curled; inflorescence greatly reduced; no fruit set; maintained via heterozygotes; resembles Cu.
pra-2 -- praeclusa-2

Plant always severely retarded, reaching 10-20 cm, unbranched, seem very thick; leaves and cotyledons small, dark green, leaves very rugose; no inflorescences seen; maintained via heterozygotes.
prc -- procumbens

Shortened internodes and leaves; rugose leaves lightly mottled yellow.
prc-2 -- procumbens-2
prc2 Older stems become prostrate.
prc-3 -- procumbens-3
prc3 Prostrate cushiony plant; elongated leaves with fewer long stalked pinnae.
pre -- pressa

Plants attain 1/3 height; habit weakly branched, semi-erect and slender; angle of leaf emergence very narrow; leaves very short, segments relatively coarse and plicate; fruit development reduced.
Pseudomonas resistance

pro -- procera

More rapid growth rate; few foliolules, larger segments entire. Tall, slender, weak plant.
prt -- protea
prt1 Smaller plant, variably reduced; pinnae keeled, pale-yellow, partly shiny, leathery.
prun -- prunoidea

All parts, particularly fruit, are elongated; traits weakly dominant.
prun-2 -- prunoidea-2
prun2 Elongated hypocotyl internodes, leaves, fruit; very lax habit.
Prx-1 n Peroxidase-1

Prx-1 5 Peroxidase-1

Prx-1 4 Peroxidase-1

Prx-1 2 Peroxidase-1

Prx-1 1 Peroxidase-1

Prx-1 3 Peroxidase-1

Prx-2 3 Peroxidase-2

Prx-2 n Peroxidase-2

Prx-2 1 Peroxidase-2

Prx-3 n Peroxidase-3

Prx-3 a1 Peroxidase-3

Prx-3 2 Peroxidase-3

Prx-3 1 Peroxidase-3

Prx-3a 1 Peroxidase-3a

Prx-4 3 Peroxidase-4

Prx-4 4 Peroxidase-4

Prx-4 23 Peroxidase-4

Prx-4 5 Peroxidase-4

Prx-4 6 Peroxidase-4

Prx-4 7 Peroxidase-4

Prx-4 9 Peroxidase-4

Prx-4 13 Peroxidase-4

Prx-4 22 Peroxidase-4

Prx-4 8 Peroxidase-4

Prx-4 14 Peroxidase-4

Prx-4 15 Peroxidase-4

Prx-4 1 Peroxidase-4

Prx-4 21 Peroxidase-4

Prx-4 10 Peroxidase-4

Prx-4 12 Peroxidase-4

Prx-4 16 Peroxidase-4

Prx-4 17 Peroxidase-4

Prx-4 18 Peroxidase-4

Prx-4 19 Peroxidase-4

Prx-4 2 Peroxidase-4

Prx-4 20 Peroxidase-4

Prx-4 11 Peroxidase-4

Prx-6 -- Peroxidase-6

Prx-7 n Peroxidase-7

Prx-7 1 Peroxidase-7

Prx-7 2 Peroxidase-7

Prx-7a n Peroxidase-7a

ps prov2 positional sterile
ps Corolla does not unfurl; pollen is not shed.
ps -- positional sterile
va Corolla does not unfurl; pollen is not shed.
ps-2 -- positional sterile-2

Positional sterile flowers; pollen is not shed. .
psa -- perspicua

Plants attain 1/2-3/4 normal size; early leaves pale light green; pinnae bent strongly upwards and more serrate, exposing undersides; distinctive yellowish light green color with exposed pale green lower surface.
pst -- persistent style

Style adnate to fruit throughout development; fruit becomes strongly beaked.
psu -- paucisurcata

Plant small, almost without branches; internodes and leaves strongly shrotened; leaf segments very small, light keeled, dull grey-green; flowers small; fruit set variably low.
pt -- petite

Small, slow-growing chlorotic plants; light green cotyledons.
pt-4 -- pseudo-triplo-4

Resembles triplo-4 in dainty habit, elongate leaf and flower parts; differs in defective flowers.
pta -- partiaria

Seedling and plant habit normal; leaf segments narrow, dainty with deeper serration and plicate, tending to be pendant; foliolules more abundant.
ptb -- protuberant

Stylar end of mature fruit protuberant.
Pto h Pseudomonas syringae pv tomato resis. hirsutum
Resistance to Pseudomonas syringae without Fenthion sensitivity
Pto 2 Pseudomonas syringae pv tomato resis.

Resistance to Pseudomonas syringae pv. tomato.
Pto -- Pseudomonas syringae pv tomato resis.

Resistance to Pseudomonas syringae pv. Tomato, race zero sensitive to insecticide Fenthion.
Pto Pto-2 Pseudomonas syringae pv tomato resis.
Pto-2 Resistance to Pseudomonas syringae pv. tomato.
Pts -- Petroselinum leaf

Leaves resemble plain parsley foliage.
ptu -- punctulata

Bushy habit with slightly shorter internodes; cotyledons and primary leaves are light green with white-green background; foliage dull light grey-green, yellow-light green virescent with fine white-yellow spots; shoot tips bluish.
pu -- pulvinata
pul Shortened internodes, stronger branching; elongate cotyledons; leaflets broad and much divided.
pu 2 pulvinata
pu2 Shortened internodes, stronger branching; elongate cotyledons; leaflets broad and much divided.
pum -- pumila

Tiny, dainty plant with few or no branches; slow growth rate.
pun -- punctata
pun1 Smaller plant, variably branched; narrow, keeled, firm pinnae, mostly finely marked in light to dark-green.
pun-2 -- punctata-2
pun2 Pinnae, filiform, thick, gray-green.
pur -- purilla

Plants much retarded, erect, and little branched; leaf edges yellow with normal green veins.
pus -- pustulata
pus1 Strongly rugose, broad, rounded pinnae; stutdy, erect bush; dark green foliage at all stages
pv -- pulviniformis

Plants reach 1/4 to 1/3 pm size, habit cushion like via controlled growth of side shoots; leaves shortened, older ones prematurely yellowing, later drying; young stems normal, developing brown. sunken spots later.
pvs -- parvistata

Plants always small, reaching 1/4 normal size; internodes and leaves shortened, leaf segments small, bright, dark green below; shoot tips bluish with anthocyanin.
px -- praecox
px1 Lax, spreading plant; small pinnae; larger, earlier fruit.
py -- pyramidalis

Cotyledons wider, hypocotyl shorter; plants reach 2/3 normal height; habit first upright, weakly branched, later becoming pyrimidal, leaves at this time are irregularly epinastic; terminal segments broader; foliage color shiny dark yellowish.
pyl -- Pyrenochaeta lycopersici resistance
py, py-1 Resistance to corky root; heterozygote intermediate, varies with environment.
r prov4 yellow flesh
r Yellow color of ripe fruit flesh.
r prov5 yellow flesh
r Yellow color of ripe fruit flesh.
r y yellow flesh reddish yellow ry Likely allele of r with reddish flesh tones in ripe fruit
r (2s) yellow flesh
r^3, r-2, r2 Yellow fruit flesh; lighter yellow flowers.
r (1s) yellow flesh
r^2 Yellow color of ripe fruit flesh.
r -- yellow flesh

Yellow color of ripe fruit flesh.
ra -- rava

Leaves down curled and gray-green; larger trichomes longer and silky.
ra 2 rava
gri Small plant; leaves small, strongly recurved dorsally, light yell to gray-green; trichomes longer.
ra-2 -- rava-2
ra2 Cotyledons heavily dotted in white; purplish hypocotyl; smaller lax plant.
ram -- ramosissima
ram1 Variably branched inflorescence; proportionately reduced all other plant parts.
ran -- ranificans
ran1 Slender shoots, pinnae, petals; variably branched inflorescence.
rc -- rolled cotyledons

Retarded growth; cotyledons uprolled.
rd -- reduced

In greenhouse plants grow normally to about 30 cm hieght; thereafter grow slowly or not at all; intorlerant of freed cuefoune.
re -- reptans

Seedlings inclined from epicotyl, mature plants recumbent; elongate internodes; less branched.
rea -- repleta
rea1 Plant denser; crowded pinnae on shorter leaves of light yellow-green.
ref -- refrenata
ref1 Strongly retarded growth; sturdy shoots; short leaves; small, narrow pinnae.
rela -- relaxata

Lax open habit; tiny plants; light green yellow tinged leaves; heterozygotes intermediate color.
rep -- repens

Strong creeping tendency even in seedling; less branching; shortened, prematurely dying leaves.
rep-2 -- repens-2

Early growth typical spl; from flowering on, all parts develop plagiotropecally, exposing fruits, which ripen earlier than normal.
rep-3 -- repens-3

Plants reach 1/3 normal size, weakly branched and plagiotropic, internodes and leaves shortened; foliage light grey-green; leaves necrose prematurely; fruits relatively large.
res -- restricta
res1 Smaller, squarrose bush; yellowish light-green, boat-shaped pinnae, purplish ventrally.
Rg-1 -- Regeneration-1

High shoot regeneration capacity from callus cultures established from root explants.
Rg-2 -- Regeneration-2

High shoot regeneration capacity.
ri -- ridged
rl Ridged leaves; retarded growth of shoots and roots.
ria -- rigidula
ria1 Smaller plant; stiff, yellowish young leaves turn dark-green later.
ria 2 rigidula
ria1^2 Smaller, stiff, compact plant; erect sideshoots; young pinnae keeled to involuted light yellow-green, dark veins, purplish below.
rig 2 rigida
pca, pca1 Small, lax, spreading plant; pinnae keeled, yellowish to light-green; early flowereing, ripeninging; heterozygote recognizable.
rig -- rigida

Plant small, rigidly erect when young; leaves yellowish, and darkening later.
rig-2 -- rigida-2

Plant small, rigidly erect when young; leaves yellowish, and darkening later.
rin -- ripening inhibitor

Fruits green at maturity, later turning bright yellow, retarded ripening.
rl -- radial cracking resistance
ra Resistance of fruit to radial cracking.
ro -- rosette

Internodes extremely short, inducing rosette plant heart; no flowers, but vistigial buds formed on older plants; height 6 inches or less; reduced branching of root system, producing a single tap root; leaflets narrower than normal.
roa -- rotundata
roa1 Small plant; short internodes, leaves; broad, rounded, rugose, light gray-green pinnae.
roa-2 -- rotundata-2
roa2 Rounded, slightly rugose pinnae; shortened leaves
rot -- rotundifolia

Short internodes; short, broad, rugose leaves.
rot-2 -- rotundifolia-2
rot2 Small plant; short internodes, sidshoots; small, broad pinnae, nearly round.
rpa -- repetita

Seedling size variable; plants weak, attaining 1/3 size; habit weakly branched, upright; leaves highly divided, each main segment equivalent to a normal leaf, early color dull grey-green, becoming normal later.
Rs -- Root suppressed

Greatly restricted or no root development.
Resistant to skin cracking

rt -- potato virus Y resistance

Resistance to potato virus Y.
rtd -- retarded dwarf

Small plant of slow growth; mature leaves drooping.
ru -- ruptilis

Leaflets narrow, keeled, dull light green, darker veined; heterozygote variably intermediate.
ru prov2 ruptilis
ru Leaflets narrow, keeled, dull light green, darker veined; heterozygote temporarily intermediate.
rust -- rustica

Dwarf plants with short internodes; leaves broad, and blunt with fewer segments.
rust-2 -- rustica-2
rust2 Small, lax bush; fewer, rounded pinnae.
rv -- reticulate virescent

Cotyledons and new leaves pale with dark veins, turning to normal green with age.
rv-2 -- reticulate virescent-2

Cotyledons light green; leaves yellowish with green veins.
rv-4 -- reticulate virescent-4

Sharply defined virescence- yellowish color of the immature leaves strongly contrasted with dark green network of the veins; seedling retarded.
rvt -- red vascular tissue

Vascular tissue deep red in roots and stems.
s -- compound inflorescence

Very compound inflorescences.

sa -- sphacelata
sa1 Small plant; light yellowish leaves, developing necrotic lesions ventrally or on pinnae tips; environmentally sensitive.
sar -- squarrulosa
sar1 Small plant, somewhat squarrose and slender branched.
scf -- scurfy

Striated, scurfy cotyledons.
scl -- seasonal chlorotic lethal

Cotyledons pale yellow, sometimes grooved; lethal in early stages except for recovery in mid summer.
sco -- scopulina

Plants reach 3/4 normal size, erect, bushy, thick coarse branches, internodes and leaves shortened; leaf segments large, broad, very wavy and rugose; fastigiate inflorescences resemble broom.
sd -- sundwarf

Very short, scarred internodes in high light intensity; heterozygotes usually intermediate.
sd-2 -- sun dwarf-2

Very short, scarred internodes in high light intensity; heterozygotes usually intermediate.
sd-3 -- sun dwarf-3

Interacts with sd-2 and br to yield birdsnest phenotype.
Se -- Septoria lycopersici resistance

Resistance to Septoria lycopersici.
sem -- semiglobosa

Smaller, compact plant; very short internodes; small dark leaves.
sem-2 -- semiglobosa-2
sem2 Smaller plant, leaves, pinnae; short internodes.
ser -- serpentina
ser1 Curved shoots, leaves inflorescences.
ser-2 -- serpentina-2

Seedlings normal; plant size almost normal; all vegetative parts, paricularly midveins and petioles irregularly elongated, result ing in twisting.
ses -- semisterilis
ses1 Smaller, erect, nearly unbranched plant; thick, yellowish, light gray-green leaves; weakly fertile in greenhouse, unchanged on normal graft stock.
sf -- solanifolia

Primary leaves entire; segments of later leaves entire and folded; filiform calyx and corolla; fruit shape often modified.
sf wl solanifolia wrinkled leaf wl, wr Primary leaves entire; segments of later leaves entire and folded; filiform calyx and corolla. Leaf margin finely wrinkled; flower parts small and distorted.
sfa -- sufflaminata
sfa1 Usually smaller, weakly branched plant; yellowish to yellow-green, involuted pinnae, purplish ventrally; slight F1 seedling heterosis.
sfa 2 sufflaminata
par Usually smaller, weakly branched plant; yellowish to yellow-green, involuted pinnae, purplish ventrally; slight F1 seedling heterosis.
sft -- single flower truss

Reduced flowers per inflorescence; indeterminate inflorescence.
sh -- sherry

Fruit flesh yellow with reddish tinge.
sha -- short anthers

Upper portion of anthers aborted; stigma protruding. Plant and seeds of low viability.
si -- sinuata

Small plant, proportionately reduced; wavy, yellow-green leaves.
sig-1 -- signal transduction-1
JL1 Deficient for production of proteinase inhibitor II in leaves following wounding
sig-2 -- signal transduction-2
JL5 Deficient for production of proteinase inhibitor II in leaves following wounding
sit -- sitiens

Smaller plant, very weak in field; very short leaves down-curled, overwilting and becoming necrotic.
Skdh-1 1 Shikimic acid dehydrogenase-1

sl 3 stamenless
sl3 Nearly stamenless, anthers much distorted; Like sl:2 except for greater anther development
sl 4 stamenless
sl4 Nearly stamenless, anthers much distorted; Like sl:2 except anthers nearly normal in winter greenhouse
sl 2 stamenless
sl2 Nearly stamenless, anthers much distorted
sl -- stamenless

Stamens abortive or absent; fruit form modified.
sl cs stamenless corollaless cs, sl^5, sl5 Corolla and androecium sepaloid, stigma irregular and protruding.
sl-2 -- stamenless-2
sl2 Stamens abortive or absent; fruit form modified. Some general features as sl except for subnormal stamens.
slx -- serrate lax leaf

Drooping, elongate, serrate leaves. Unknown variety from Romania.
Sm -- Stemphyllium resistance

Resistance to Stemphylium, solani.
sms -- small seed

Endosperm almost absent; testa thin and transparent; reduced germination.
sn -- singed

Epidermal hairs smaller; trichomes crooked, curved and otherwise distorted.
snt -- Snout
sn Persistant style (beaked fruit)
so -- soluta

Plant size normal, habit loose; pinnae narrow, pointed, deeply serrate, margins slightly elevated; similar to prunoidea but with normal fruits.
Sod-1 1 Superoxide dismutase-1

Sod-2 1 Superoxide dismutase-2

sp D self-pruning Dominant
Dominant allele; severe determinate growth habit; homozygous lethal. Radiation induced in Stupicka polni ranna.
sp prov2 self-pruning

Plant habit determinate.
sp + self-pruning

Indeterminate growth habit.
sp -- self-pruning

Plant habit determinate.
spa -- sparsa

Plant size reduced; leaves emerge yellow-green, becoming blotched whitish yellow-green.
spe -- splendida
spe1 Small plant; shiny, yellowish foliage; older leaves darker than younger ones.
Spf -- Superpuff

Extremely puffy, hollow locules and bell pepper shaped fruit.
Spf-2 -- Superpuff-2

Extremely puffy, hollow locules and bell pepper shaped fruit.
sph -- sphaerica

Smaller, spherical bush; short internodes; older leaves firm, dark green, and shiny.
Spi 2 Sympodial index

Sympodial index-two nodes. An average of 2 leaves between successive inflorescences.
spl -- splendens
spl1 Shiny, yellowish to light green, boat-shaped, rugose pinnae.
spl-2 -- splendens-2
spl2 Lax, flat bush; short leaves; small, narrow, pointed, keeled, crowded pinnae.
spl-3 -- splendens-3
spl3 Flat, dense bush; narrow, pointed, keeled pinnae.
spl-4 -- splendens-4

Distinctive seedling resembling spl; interveinal areas brilliant yellow; leaf margins tend to roll inward especially near growing point.
squa -- squarrosa

Small, very weak plants of squarrose habit, few branches; small, pale gray-green leaves; heterozygote intermediate in color and size.
squa-2 -- spuarrosa-2
squa2 Narrow, light green cotyledons; diminutive plant, leaves, and pinnae.
sr -- slender stem
sm Stem slender, stiff, upright, with few branches; leaves elongate, blue-green, with few segments.
sru -- subristica

Growth always retarded, plants reaching half normal size; cotyledons short, rounded and blunt; heavily branched, internodes shortened; leaf segments broad, ovate, slightly rugose, and dark green.
ss -- spongy seed

Smoothe but spongy seed surface.
st -- sterile

Completely sterile; plants later become purple.
sta -- stabilis

Seedlings normal with marked later reduction in growth, plants reaching 1/3 pm size; habit small round bush, internodes being proportionately much more foreshortened than leaves and strongly branched.
stam -- staminodea

Staminoid petals; anther tips free and more slender.
ste -- sterilis

Smaller plant; numerous, rugose, smaller pinnae; many flower buds, often aborting; weak fruit and seed set.
str -- striata
str1 Pinnae with downward rolled margins and finely dotted in yellow early, turning green later except at base.
stri -- stricta

Rigid upright growth; broader blistered leaves.
stri-2 -- stricta-2
stri2 Larger plant, leaves, pinnae; stocky stems.
stri-3 -- stricta-3
stri3 Larger, dark to strong green pinnae, margins rolled upwards.
stri-4 -- stricta-4
stri4 Large, stocky bush; rounded, rugose, moderately dark green pinnae.
stu -- stunted

Very slow growth; strong anthocyanin.
su 3 suffulta
di Rachis very short, leaf color yellowish gray-green; stems white, streaked green.
su 2 suffulta
exa Leaves deeply veined, yellow-tinged; fruit cluster upright; strong anthocyanin.
su -- suffulta

Leaves deeply veined, yellow-tinged; fruit cluster upright; strong anthocyanin.
su ni suffulta nitida di^ni, ni Leaves long-petioled with dainty, deeply cut segments
sua -- suffusa

Small, broad plant; paler leaves, yellowing prematurely.
sua-2 -- suffusa-2
sua2 Small, keeled, light green pinnae; branched inflorescences.
sub -- subtilis

Tiny plant with short internodes, fastigiate habit; narrow acute leaf segments.
suc -- succedanea

Small, weakly branched, erect plant; short, firm leaves, yellowing prematurely.
sucr -- sucrose accumulator
TIV1 Accumulates predominantly sucrose in mature fruit, rather than glucose and fructose.
suf -- sufflava

Uniform light green color.
suf-2 -- sufflava-2
suf2 Larger, lighter cotyledons; small plant and leaves.
sul -- suckerless

reduced side-shoot formation; often a few basal shoots are produced. Inflor. With 1 - 2 flowers only.
sulf pura1-10 sulfurea

Cotyledons greenish yellow, turning pale yellow; true leaves yellow; lethal unless grafted on normal stock.
sulf vag 1-3 sulfurea

Leaves variegated green-white; some homozygotes lethal unless grafted
sun -- Sun1642 fruit shape

Elongated fruit shape, from Sun1642.
suo -- subrotunda
suo1 Many slender side shoots; very small, irregularly rounded, dark gray-green pinnae.
sup -- superba

Seedlings normal, plants later becoming strongly upright and elongated; strong mainstem and petioles; leaf sefments large, coarse, and blistered, dark, and shiny.
sur-2 -- suberosa-2
sur2 Necrotic flecks on the deformed whitish cotyledons and older pinnae.
Sus -- Subsistens

Growth variably suppressed; few or no branches; varied meiotic abnormalities; homozygous lethal.
Sw b Spotted wilt resistance

Resistance to spotted wilt strain TB2.
Sw a Spotted wilt resistance

Reisistance to strains spotted wilt TB, N, and R.
sw-2 -- spotted wilt resistance-2
sw2 Resistance to Sw strains TB:1, N:2, R:2, and R:3.
sw-3 -- spotted wilt resistance-3
sw3 Reistance to SW strains R:2,R:3, and M:2.
sw-4 -- spotted wilt resistance-4
sw4 Resistance to Sw strains M:1 and M:2.
Sw-5 -- Spotted wilt resistance-5

Resistance to to Tomato Spotted Wilt Virus. From L. peruvianum.
Sx -- Sepal extender
sx Elongated sepals; heterozygote intermediate.
sy -- sunny
ye Cotyledons green bleaching to yellow; yellow virescent.
syv -- spotted yellow virescent

Bright yellow virescent; fine whitish speckling on mature leaves; can turn necrotic.
t -- tangerine

Fruit flesh and stamens orange colored.
t 2 tangerine
t1^2 Yellowish growing point; light green foliage; resembles t in flower an fruit color
t v tangerine virescent
Furit and flower color typical of t; irregular yellowing near growing point.
ta -- tarda

Smaller, dainty plant; leaves reduced, glossy, gray-green.
tab -- tabescens

Normal seedling growth followed by severe stunting; leaves irregular, yellow-green, violet veined with necrotic specklings.
tc -- turbinate corolla

Plant smaller; petal tips turbinate.
te -- terminata
te1 Determinate growth; fasciated inflorescences, flowers, and fruit.
tem -- tempestiva
tem1 Small plant, short internodes, leaf midribs; involuted pinnae, lighter with darker veins.
ten -- tenuis

Very slow growth; leaves light green, reticulated with whitish yellow veins.
tf 2 trifoliate
tri Leaf usually with only three segments, petiole elongate.
tf -- trifoliate
ct, tri Leaf usually with only three segments, petiole elongate; reduced fertility.
ti -- tiny plant

Plant about 1/4 normal size; stiff leaves and branches.
tl -- thiaminless

Cotyledons normal; leaves small, yellow, developing green veins, and dying prematurely; lethal; viable and normal if fed thiamine.
Tm -- Tobacco mosaic virus resistance

Moderate level resistance to TMV.
Tm-2 -- Tobacco mosaic virus resistance-2
Tm2 High-level resistance to many strains of tobacco mosaic virus.
Tm-2 a Tobacco mosaic virus resistance-2
Tm-2^2 High-level resistance to many strains of tobacco mosaic virus.
tmf -- terminating flower

First inflorescence with single, abnormal flower; lateral branches have normal inflorescences.
tn -- tenera

Plants attain 1/3 typical size; habit erect, rarely branching; stems slender; foliage color dark green early, becoming irregular with variable anthocyanin content later.
Tor -- Tortilis

Coiled or downrolled cotyledons, leaves, or petals.
tp -- tripinnate leaf

Leaves tripinnately compound; retarded growth.
Tpi-2 1 Triosephosphate isomerase-2

tr -- truncata
tr1 Smaller, weak plant; short, gray-green leaves, yellowing prematurely.
tr-2 -- truncata-2
tr2 Blue-green seedlings; smaller plant, leaves, pinnae; older leaves dying prematurely.
tri 1 temporarily red light insensitive

Insensitive to red light for up to 2 days; hypocotyls elongated and cotyledons reduced compared to wild type.
trs -- tristis

Seedlings 1/4 normal size; leaves narrow, plicate and strongly pendant.
tu -- tumida

Plants reach 1/3-2/3 normal size, fewer branches; main stem bent at nodes, which are swollen; cotyledons and leaves deformed, wavy, blistered; petals and sepals narrow, wavy, and anther often fringed.
tur -- turgida

Growth retarded at all stages; branching strong, internodes shortened, dense bush habit; leaf segments long petioled, light yellowish margins; flowers smaller.
Trialeurodes vaporariorum resistance-1

Trialeurodes vaporariorum resistance-2

Ty-1 -- TYLCV resistance

Tolerant of TYLC virus.
u Fs uniform ripening fruit striped Fs Dominant allele; dark shoulder; dark green radial striped opposite each locule of highly pigmented unripe fruit
u -- uniform ripening
u1 Unripe fruits of uniform light green color, lacking normal darker shoulder.
u J uniform ripening Jubilee
Very light green shoulder; resembles u:G
u G uniform ripening Galapagos
Resembles u except for partial dominance.
u + uniform ripening

Presence of green shoulder on immature fruit.
ub -- umbraculiformis

Plants attain 2/3 typical size; habit erect; leaves coarser with flatter pinnae; young leaves convex.
uf -- uniflora

Inflorescence with single flower.
ug -- uniform gray-green
u2 As of u except for grayish color.
ul -- upright leaf

Upright leaf.
um -- umbrosa

Mature leaves darker green, wilted appearance; later growth stunted; reduced root growth.
um-2 -- umbrosa-2
um2 Small, lax bush; smaller pinnae, dark green to dull gray-green.
um-3 -- umbrosa-3
um3 Broad, crowded, dark green pinnae; small leaves, short internodes; small, erect bush.
un -- unijuga
un1 Very small plant; narrow pinnae, cotyledons; fewer flowers.
und -- undulatifolia

Habit flatter than normal; cotyledons and primary leaves are narrow, acuminate, and margins are deeply notched; leaf segments small, keeled, wavy, and dull light grey-green.
uni -- unicaulis

Weak, thin, upright, growth with great suppression of branching.
uni (1s) unicaulis
suc-2, suc2, uni^2 Purplish grey green seedlings; leaves and pinnae smaller, short, rounded,
uni-2 -- unicaulis-2
uni2 Narrow, slightly deformed cotyledons; very small pinnae, leaves, and plant; slow-growing.
uni-3 -- unicaulis-3
uni3 Strongly retarded growth and branching; pinnae 1/3 normal size; strong anthocyanin.
uni-4 -- unicaulis-4
uni4 Retarded growth and branching; very small, keeled pinnae.
up -- upright pedicel

Pedicels straight, oriented upward.
upg -- upright growth

upright growth habit.
v -- virescent

White virescent, whtite leaf marquis; stunted growth.
v-2 -- virescent-2
v2 New leaves pale under greenhouse conditions; frequently undistinguishable in field.
v-3 -- virescent-3
V3 Leaves small, green and white patterned virescent.
va virg varia virgata
Like va:dec except older leaves retain yellow in region of veins.
va dec varia decolorata
Bright yellow-green virescent.
va-2 -- varia-2
va2 Smaller plant; at flowering whitish yellow-green zones move from pinnae tips to bases.
val -- valida
val1 Sturdy shoots; firm, keeled, rugose, light green pinnae, blue-green below in the shoots.
var -- variabilis

Leaves light green, yellow-green virescent; slightly smaller plant with shorter internodes.
var-2 -- variabilis-2
var2 Smaller plant; at flowering shiny yellow-green growth zones, later turning light green.
Ve -- Verticillium resistance

Resistance to verticillium.
ven -- venosa

Variably reduced growth; tiny folded leaves, whitish yellow with green veins.
ver -- versicolor
yv-4, ver1 Younger leaves with fine mottling of yellow; green veins.
ver-2 -- versicolor-2
ver2 Smaller plant, leaves; light green pinnae weakly flecked in white, margins strongly uprolled.
ves -- versiformis
ves1 Pinnae pointed, keeled, bright green; unripe fruit darker.
ves-2 -- versiformis-2
vf Small, compact plant; dainty, shiny, yellowish pinnae, wavy margin.
vg -- vegetative

Greatly deformed and usually functionless flowers.
vga -- virgulta
vga1 Smaller plant; short internodes; dull-green foliage, light yellow-green growing points.
vga-2 -- virgulta-2
vga2 Leaves yellowish light-green, green margins and veins.
vi -- villous

Stems very hairy.
vio -- violacea

Heavy anthocyanin on stems and veins; dull light green leaf color.
vio-2 -- violacea-2
vio2 Light green foliage; shoot tips, petioles, leaf veins with strong anthocyanin.
vir -- viridis

Early leaves dark green, third order leaf divisions.

vlg -- virescent light green

Seedlings 1/3 normal size with light green virescence, leaves broad,; mature plants normal size, leaves light green.
vms -- variable male-sterile

Abortive anthers, reduced corolla high temperature; normal otherwise.
vo -- virescent orange

Yellow virescent; mature leaves pale bluish green; fruit flesh orange, redder in outer walls.
vra -- viridula
vra1 Small plant; boat-shaped, dull light-green pinnae, wavy margin.
vra-2 -- viridula-2
vra2 Yellowish light green leaves, darken after onset of flowering; growth retardation.
vrd -- viroid

Scalloped cotyledons; highly distorted leaves with white speckled chlorosis; extreme environmental sensitivity, winter expression best.
vt -- vieta

Plants attain 1/3-1/2 size; habit erect with robust main stem and few branches; young pinnae slightly undulated; foliage color normal except yellow-green virescent.
vta -- virgatula

Plants reach 1/4-1/2 normal size; cotyledons narrow; plant color dull light grey-green; stems blue with anthocyanin; young leaf segments rugose and striped yellow-olive.
w -- wiry

Leaves progressively reduced to slender, thread-like elements; dwarfed plants.
w-2 -- wiry-2
w2 More extreme strap shaped leaves, petals, and sepals than w:1, plant sterile.
w-3 -- wiry-3
w3, w2 Resembles w1, except ovary more syncarpous.
w-4 -- wiry-4
w4 Similar to previously described wiry mutants; leaves progressively more reduced.
w-5 -- wiry-5
w5 Resembles previously described wiry mutants.
w-6 -- wiry-6

In splendens stock. Cotyledons with shorter lamina, longer petiole, bent downward, grey-green; primary leaf almost string-like, later also strongly epinastic; next leaves with sac-like terminal segment on hair-like petiole; later leaves with 1-5 lateral segments, often partly fused; corrolla segments narrow; plants 1/3 to 1/2 normal size; produces seed in field. Typical w phenotype can be selfed.
w-7 -- wiry-7

Smilar to w-6 but with more extreme expression and less fruit set; not allelic to w-6.
w-8 -- wiry-8

Similar to w-7.
Wa -- White anthers

Anthers white or cream-colored.
wd -- wilty dwarf

Grayish-green, droopy leaves; stunted plants; leaves droop when drought stressed.
wf -- white flower

Corolla color is buff, light tan, or white.
Wlt -- Wilty

Leaves overwilt when drought stressed. Wilting under field or greenhouse conditions; marginal leaf narcrosis.
Wo mz Wooly

All parts densely pubescent; homozygous lethal.
Wo v Wooly

Denser hairiness than Wo; homozygous lethal.
Wo m Wooly

Pelage on all parts wooly but less than Wo; homozygous viable; heterozygote intermediate.
Wo -- Wooly

All parts densely pubescent; homozygous lethal.
wt -- wilty

Leaf margins curl adaxially
wt-2 -- wilty-2
wt2 Leaf margin rolled upwards at all stages.
wv -- white virescent

Plant retarded; cotyledons whitish yellow; leaves become white virescent, always with white speckled.
wv-2 -- white virescent-2

White, green-veined virescent, plant retarded.
wv-3 -- white virescent-3

Seedling 1/20 normal size, cotyledons, white or greenish white, virescent.

x -- gametophytic factor

Ineffective microgametes associated with I.
x-2 -- gametophytic factor-2

Altered transmission rate associated with c.
Xa -- Xanthophyllic

Leaves yellow; growth retarded; homozygous inviable.
Xa-2 -- Xanthophyllic-2
Xa2, A Resembles Xa.
Xa-3 -- Xanthophyllic-3
Xa3 Darker green than Xa.
xan -- xantha
xan1 Yellow cotyledons; lethal.
xan-2 -- xantha-2
xan2 Yellow cotyledons; lethal.
xan-3 -- xantha-3
xan3 Yellow cotyledons; lethal.
xan-4 -- xantha-4
xan4 Yellow cotyledons; lethal.
xan-5 -- xantha-5
xan5 Yellow cotyledons; survives under optimal conditions.
Xanthomonas campestris resistance

y -- colorless fruit epidermis

Fruit epidermis lacks pigmentation.
yc -- yellow calyx

Calyx turns yellow when fruit ripens.
yg -- yellow-green
yg1 All foliage yellow-green, vireescent.
yg slo yellow-green

yg-2 aud yellow-green-2
yg-2^r, aud All foliar parts bright yellow under all conditions.
yg-2 -- yellow-green-2
yc, yg282, yg2 Foliage chlorotic yellow-green under all conditions; long hypocotyls.
yg-3 -- yellow-green-3
yg3, yg330, ye Resembles yg4 with leaves appearing ragged.
yg-4 -- yellow-green-4
yg4, yl, yg333 Foliage chlorotic whitish yellow-green under all conditions; leaves more deeply serrate, edges curled.
yg-5 -- yellow-green-5
yw, yg388, yg5 Very yellow cotyledons and leaves; slow growing; good plant viability.
yg-7 -- yellow-green-7

Leaves light yellow-green, dotted in early growth; flowers normal; pollen inviable; growth weak. Radiation induced.
yg-8 -- yellow-green-8

Leaves bright yellow at all stages.
yg-9 -- yellow-green-9

Pronounced yellow-green leaves, becoming somewhat darker with age.
ys -- yellow seedlings

Lethal in early stage.
yv ms yellow virescent masculosterilis
Male-sterile allele of yv; seedling yellow becoming yellow green (field) or light green (green house).
yv mut yellow virescent mutabilis
Male sterile variegated allele of yv; green color frequently mutating to yv: ms
yv 3 yellow virescent
vel Light yellow-green leaves, paler at growing point, which has velvety appearance.
yv 2 yellow virescent
vel^2, vel1^2 Smaller, irregular bush; short internodes; soft yellow, velvety growing points shading to normal green below; smaller than vel.
yv -- yellow virescent

Pale yellow-green; virescent foliage.
yv-2 -- yellow virescent-2

Smaller, irregular bush; short internodes; soft yellow, velvety growing points shading to normal green below; smaller than vel. C. R.
yv-4 -- yellow virescent-4

Narrow, yellow green leaflets, dark green veins, becoming irregular, grey green.